CRLSSI
SMITHSONIAN [INS TEPULTLCON
MUSEUM OF NATURAL HISTORY
UNITED STATES NATIONAL MUSEUM BULLETIN 280
A Revision of the Peoritinae and Anerastiinae (Auctorum) of America North of Mexico
(Lepidoptera: Pyralidae)
JAY C. SHAFFER Visiting Research Associate Smithsonian Institution
SMITHSONIAN INSTITUTION PRESS
WASHINGTON, D.C. 1968
Publications of the United States National Museum
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Contents
Page
ME TOGIUECtOMMe es skeet = oe ee hes oe Sa a ae ee ete Se 1 eer n ane NC OUS! ci 8 os roe Ee ee ee EE 2 NEKO WCCO OMEN UGS = =o ee nebo 2 aa Mee, eee ye SE eee 4 emo waeeeune Witerauures o-oo en ee he eh Se 5 SURE RpesSSNTE ota AU Pee eas Sees at io a Shes Rel 2) Pe Oe 6 RECENT VE. CORMM AGL. £20 ae" eto OS Oe Sok a he iastl TO0 eo ee ae as fe 8 iG GmiEaeleCOrid AG ON OG 2s ct Sen ed Da Ra ad ee es 9 Te ensngl pel. (ALA POnOL) — 8 2... se ee ee 13
PU THOTCCLUA HAC ON OG: 2 mn os eas | el ee ee 14
tel adellan (Zeller) en sets ope ee eae oe eg 16 FIMODECCHCY CELMIBG es = 2 2s = Be Se a Se a ee ie 18 Perar relia SHalebe Sees Ses ee ot oa a ee ey oe ee 20
Er astrelig: Wiagongt) 222s" wn eg eee A ela ee ee ee 21
FQ CIIGL CLL G™ (UNS G) aes a OS el oy he ee ed 22 PaevOseOlnrrella, (RS PONO) 2 ~ se 2 eo ee 24 PRPIONMSLON THA CE soo ee eo Ae nae: Mb cece Rate Sys ees ates 26
Pas ntnTeeUe (Cl) Vary ache tie cee 2 ees eed 8 Leak We 27 PUNOLO DON ELENGN OLY VAT ete 5.70 ieee) es SE i Pa 2 St a ys rg 28 NOD DROTUILEL Gs (WAIT) tae SOA. 8 ee red ae Py ee 29
em LEICOSICHE (CREE ONTO eee et a i Sr a a pes 36 SETTER TRIS J A a a et a Ng a oe ees OM Jp rg TALIS MEST DEN 12) i ces ce is a a Pe gE ier ee eens Meese 38 iBertineAriVvacg planers Get wees My 9k ea ee ae A Ae 39 miesbimella | Groups secu as eae L) yO vem Steed Saeed Chas et ee S28 40 Penne UO yar) ete ee ban Pe Se Gee ee See es 41
PAPO LITVCLL ORS ATT OR eee = om at en eel ST ees ela eer es oy es en Seep es ees 42 Mibu Snrecne Grr OUEN ss. eee ek ee ee ee ee ee 42 PAOSICECLLE TUR POROU) eo ee ee Seen eee eee ee Oe ee ee ee 43 CASTE TY CLI 2s SM ORS SEA Se ah a eS Tae Ee oon A 44 she vAlbidelie Groupe | 25 hehe yah he Cates hate ah a A ee 45
PAPE LUTCELL@ A CEMUISE) rien Spee es ee a ek es We ee oe 46
PAT OT LOUIS ALT Ci hae a he a Be pe gS a eh 46 PRE PAT DIOORDCHA LOU Piet te ns ee en) Mee a Fe 47
APRIL EEFECO SEE TEE (AGrE OSS OCC) ee re Nees eee eee eee ee I Se 47 CRCHUIBSATESCOSO TIMUIGG =~ 02 oe eee Ree eee es eS 48 ARTO LaTeOSellai (Zeller) eee: Spy ssh Pee Wi gel es Ae eee eres 49 ORiseL omosassa: HUIS by jo So ons ot ee EP Fe a es ee 50 SEI CLR ELVIN Gs 28 2 A See ee eek ee I fs oe oe oe 51
PPAR AD CREE PUES DLR OTS os a a yt ee De eee EE 53
EHR TCC TLCLEULC SIN PULL © Eis ee ep eae eee Se a es re ee 54 Geniisteynoce ouniiera. ©. blll tie ee ee aes Se oS 54 piLOSCeLLON (UUIS EERE eee et yaa ce ae een Epes ie eer 55 CCRMISIG OU GUE AP ONO Ge ooo) Megat Se i oS ee ee 56 Gepstictelas (Pram pEOn) o> 5/52 2) 2 eS Bere ee ea 56
VI CONTENTS
Page
Sublamuily Phycitingae: ds. 2 2. a. fe eel ee ee eee ee rs ee 57 Genus Anerasiza Titbner. 26:22 at a ee 58 Are LOCeL GG TANG ICT ees a eet ee ep rer 59 Gemus) Coenochroa Magonot. 2-65 ae ee ee eee 60 Cmcalioniiela havonotese 22422 ease ee er eee 61 CAI bella ann st) ee ee a ee ee Le 65
C. bipunctella. (Barnes and: MceDunnough) ==+---_ 2-22-25 2= 67 Genus Bandera -Ragonots =e = aes es ee eee ee 68 B.bunotelian(Zeller)t44-< 555 ees et agen Cee 69
Ba cupvdrrellay Ta Sts ep a ee ee vel Bevrgmella-Dyars sean. SS eke ee ee 73 Genus Wakulla’ Shaffer. 2220. 22-24 See a re ee 74 W. carneella (Barnes:and MeDunnough). = 2225 eet wo Genus Tampa: Ragonot-2s222.c2 Soe 2 eS ee eee 75 Lf. dimedtatella Ragonot.. 2222 a ee ee a ee 76 Genus Barberia Dyars. 2222 ee ee ee 78 B. afinitella Dyariow ioe noes ee cn A eee 78 Genus/Hagonolia Grote: 22a ee ee eee 79 Redotaliss(Hulst) 222 0. sa i ek 80 Gens Martia>Ragon0te.2522.24 5400 bo et ee ee 83 M.arizonella Ragonoto.s22. Sa seo es 2 a. | ed ee 84 Synonymicall List of Species \Considered- =. 2.2 Se ee 85 Unplaced: Genera: and'Species 2 us a: oe 87 Initeravure Cited iis? 24 2. Shee. ee eee ed eee aie 90
Bxplanation of Figurés: 2-2) as2222 2 Yel Le OR ee 93
Introduction
This study began as a revision of the North American Anerastiinae, largely in the sense of the McDunnough (1939) checklist. In keeping with the original goals of the investigation, this paper treats all species formerly placed under that subfamily name, although Anerastia and several other genera are herein transferred to the Phycitinae, and Peoria is taken as the nominal genus of the subfamily. The transferred genera have not previously been treated in detail and so are discussed in this paper. The following introductory sections deal with the litera- ture and history of the classification of all the genera formerly placed in the Anerastiinae.
The species of moths which previously constituted the subfamily Anerastiinae are a mixture of various pyralid elements. Members of this worldwide group are generally uncommon, rather inconspicuous insects of little or no economic importance, and perhaps largely for these reasons have received relatively little attention from taxonomists.
Much of the work which has been done on the North American Anerastiinae appeared during the last 15 years of the 19th century and is superficial by current standards. The only world monograph of the group is that of Ragonot (1901). Because existing works on the anerastiines have not been based on the detailed examination of species which is so necessary to a natural taxonomic system, a number of areas remain as major problems.
Identification of species has been difficult, even for specialists, be- cause the descriptions and illustrations are inadequate. Most of the original descriptions occupy but a few lines of text and are almost useless for identifying the species. Illustrations of the adults are nearly absent from the literature with the exception of the Ragonot revision, a rare publication not easily available to most workers. The genitalia, one of the most taxonomically useful structures, are nowhere illus- trated for North American species and are described in only a few instances. In addition most past studies were made from only one or a few specimens so that no account of individual and geographic varia- tion was possible. Thus, reliable identifications can be made only by comparison with the type specimens. This is difficult because the types are scattered in various museums throughout the United States and Europe.
Placement of species into genera has been on the basis of usually one or a few superficial characters of wings, antennae, and palpi,
1
a U.S. NATIONAL MUSEUM BULLETIN 280
without emphasis on the sum of these characters and without examina- tion of the genitalia. As a result most species have been misplaced, often associated in Old World genera to which they have no close affinities.
The Anerastiinae and Phycitinae have long been considered closely related within the Pyralidae and were often collectively recognized as the family Phycitidae. The two groups show close similarity in wing venation, notably the loss of vein R; in both groups. They also have similar palpi and the frenulum reduced to a single bristle in the females.
Separation of the Anerastiinae from the Phycitinae has been made on the basis of a single reduction character, loss or extreme reduction of the tongue, hence the supposed monophyletic nature of the group has been highly questionable. The results of the present investigation indicate that in North America eight genera properly belong with the phycitines and seven genera, forming a natural group, are quite dis- tinctly separate from the Phycitinae. The latter of these two groups has been given subfamily status as the Peoriinae, a subfamily name proposed by Hulst.
The present study is preliminary; essentially all that is known of the group is based on studies of the adults; the immature stages and the biology of the living insects are almost wholly unknown. Food plant records are available for only two species of peoriines, both grass feeders. Peoria bipartitella is reported to feed on Panicum lanuginosum, and Peoria tetradella on Elymus canadensis as a borer in the stem. The life history of Anerastia lotella, a Holarctic species, is well described in the literature, notably by Buckler (1901). The species feeds on a number of grasses including Ammophila arenaria, Festuca ovina, Secale cereale, and Azra species. It is hoped that this paper will encourage taxonomic and biological studies on the Peoriinae throughout their worldwide range.
Materials and Methods
SPECIMENS EXAMINED.—The U.S. National Museum and the Canadian National collection have provided most of the specimens used in this study. A complete listing of individuals and institutions from which material was borrowed is given below along with the abbreviations used in the text.
ABK Collection of Alexander B. Klots, New York, N.Y.
AMNH American Museum of Natural History, New York, N.Y. ANS Academy of Natural Sciences, Philadelphia, Pa.
BM Collection of Bryant Mather, Jackson, Miss.
BMNH British Museum (Natural History), London, England BPI State Bureau of Plant Industry, Harrisburg, Pa.
CM Carnegie Museum, Pittsburgh, Pa.
CNC Canadian National Collection, Ottawa, Ontario
REVISION OF PEORIINAE AND ANERASTIINAE a
CNHM Chicago Natural History Museum, Chicago, Ill. CPK Collection of Charles P. Kimball, Sarasota, Fla.
CU Cornell University, Ithaca, N.Y. INHS Illinois State Natural History Survey, Urbana, III. JCS Collection of the author, Shamokin Dam, Pa.
JGF Collection of John G. Franclemont, Ithaca, N.Y. KSU Kansas State University, Manhattan, Kans. LACM Los Angeles County Museum, Los Angeles, Calif. MOG Collection of Murray O. Glenn, Henry, II}.
UCB University of California, Berkeley, Calif.
UK University of Kansas, Lawrence, Kans.
UM University of Minnesota, St. Paul, Minn. USNM U.S. National Museum, Washington, D.C.
TECHNIQUES.—Preparatory to examining the genitalia the abdomen was broken away from the thorax and macerated in cold 10 percent potassium hydroxide (KOH) solution for from 4 to 8 hours, then dissected in 50 percent alcohol. The abdomen was carefully torn open with jewelers’ forceps along the entire right pleural area, enabling the cuticle to be mounted flat with the outer surface upward. The genitalia were separated by carefully tearing along the membrane between segments eight and nine. After the removal of loose scales and KOH, the male genitalia were usually stained with mercurochrome (0.3 percent aqueous solution), or occasionally with chlorazol black or acid fuchsin to improve rendition of various structures. The aedeagus was removed, and the valves were spread and held open by a small rectangular section of microscope slide placed over the genitalia. The genitalia were held in this position in 95 percent alcohol until hardened, then transferred successively to clove oil and xylene for about 15 minutes each, prior to mounting in Canada balsam on a microscope slide.
Female genitalia were put through the same series of chemicals as those of the males, but were usually stained with chlorazol black or mercurochrome.
In order to allow observation from any desired angle the genitalia of at least one specimen of each sex of each species were transferred from xylene into small (5 x 10 mm) genitalia vials, along with a drop of Cargille’s type A microscope immersion oil; the males were left unspread. These specimens were eventually washed in xylene and mounted in balsam; before mounting they proved very useful in preparing descriptions because it was possible to study the relation- ships of the various structures.
Attempts to spread the male genitalia of the Peoriinae in the manner described above caused considerable distortion; thus the method had to be modified. If one will visualize the external male genitalia as the aedeagus and a surrounding cylinder, and then the cylinder as severed lengthwise, the genitalia can be ‘‘unrolled” and
4 U.S. NATIONAL MUSEUM BULLETIN 280
flattened. In practice this was achieved by carefully separating the right valve from the vinculum and tearing the diaphragma so that the juxta remains adjacent to the right valve. The method was highly satisfactory as all the genitalic structures then lay flat in one plane. The genitalia were held flat, run through the alcohol, clove oil, xylene series, and mounted as explained above. Illustrations of the peoriine species and several other species were made with the male genitalia prepared in this manner.
Trrms.—The Comstock-Needham system has been used in naming the wing veins (fig. 50). I have followed Kornerup and Wanscher (1962) for color terminology, viewing both the color plates and the specimen by incandescent illumination. Physiographic distribution terminology follows Lobeck (1948). Unless otherwise indicated, the phrase “North America” is taken to mean America north of Mexico.
Acknowledgments
The investigation was carried out under the direction of Prof. John G. Franclemont, to whom I wish to express my gratitude for his kind advice during the course of the project and for his helpful criticisms of the manuscript. A special note of appreciation is due Mr. Paul E. S. Whalley, who checked several type specimens in the British Museum (Natural History) against my descriptions and photo- graphs; to Dr. Pierre E. L. Viette, who made available many of the Ragonot types in the Paris Museum; to Dr. Alexander B. Klots for his numerous helpful suggestions; to Dr. J. F. Gates Clarke, who sug- gested and encouraged my work on this interesting problem, and to Dr. H. E. Moore, Jr. and Dr. W. D. Duckworth, who offered useful comments on the manuscript.
I wish to thank the Allied Chemical Corporation for a contribution toward the cost of preparing the manuscript and for a grant which enabled me to spend one month collecting specimens at the Archbold Biological Station, Lake Placid, Fla. I am also indebted to the pub- lisher of the Goode Base Map Series, Department of Geography, The University of Chicago, for granting permission to reproduce copyrighted material used in several of the plates. Maps 1 through 6 and Map 12 are based on Goode Base Map No. 202, copyright by the University of Chicago.
Finally, the investigation would not have been possible without the kind cooperation of numerous individuals and institutions who loaned specimens for my studies. I extend my appreciation to Dr. George W. Byers, University of Kansas; Mr. Harry K. Clench, Carnegie Museum; Mr. Hugh B. Cunningham, Illinois Natural History Survey; Dr. Donald R. Davis, U.S. National Museum; Dr. Howard E. Evans,
REVISION OF PEORIINAE AND ANERASTIINAE 5
Museum of Comparative Zoology, Harvard University; Dr. John G. Franclemont, personal collection; Mr. Murray O. Glenn, personal collection; the late Dr. Harold J. Grant, Jr., Philadelphia Academy of Natural Sciences; Mr. Charles P. Kimball, personal collection; Mr. Norman Marston, Kansas State University; Mr. Lloyd M. Martin, Los Angeles County Museum; Mr. Bryant Mather, personal collection; Dr. Eugene Munroe, Canadian National collection; Dr. L. L. Pechuman, Cornell University; Dr. Jerry Powell, University of California, Berkeley; Dr. Frederick H. Rindge, American Museum of Natural History; Mr. George B. Sleesman, Pennsylvania State Bureau of Plant Industry; Dr. Frederick W. Stehr (now at Michigan State University), University of Minnesota; Mr. Paul E. S. Whalley, British Museum (Natural History); and Mr. Alex Wyatt, Chicago Natural History Museum.
Review of the Literature
The genus Anerastia Hiibner [1826] originally comprised four species; three of these were transferred to Hypochalcia Hiibner by Zeller (1848), and the one remaining species, lotella Hiibner [1810- 1813], was assumed to be the type of the genus although it was not explicitly designated as such until 1886 by Moore. In the original publication of lotella, Hiibner gives only a watercolor figure, no description. Genitalia of both sexes were figured and described briefly by Pierce (1938).
Walker (1866) originally described approximella, the same species later being named haematica by Zeller (1872), and roseatella by Packard (1873). Zeller’s paper included descriptions of three other new species of North American Anerastiinae.
In 1886 Ragonot established the family Phycitidae, based on Phycita Curtis, and divided it into the subfamilies Phycitinae and Anerastiinae based upon the ‘‘absence, or nearly so, of the tongue” in the latter group. He states his intention to prepare a monograph of the Phycitidae and Galleridae of the world and gives a review of previous works on these insects.
The publication of Hulst’s ‘Descriptions of New Pyralidae” (1886) prompted Ragonot to publish new species of his own in advance of the projected monograph. The appearance of the Hulst papers at this time was in some respects unfortunate, for in the next several years both men published many new specific and generic names without seeing each other’s types, thereby creating many synonyms. During 1887-89 Ragonot published 16 species and 15 genera, and Hulst published 7 species and 4 genera listed as new for the North American Anerastiinae.
6 U.S. NATIONAL MUSEUM BULLETIN 280
Grote (1888) regarded “. .. the Phycidae or Phyciinae as a sub-family of the Pyralidae; the M. Ragonot’s Anerastinae [sic] as merely a tribal division of the sub-family.”
The first paper summarizing knowledge of the Anerastiinae was Hulst’s (1890) ‘Revision of American Phycitidae.”” Therein he divided the family into the subfamilies Phycitinae and Peoriinae, the latter group largely corresponding to Ragonot’s Anerastiinae, but excluding Anerastia and two other genera. The division was rather auspiciously made on the basis of the male genitalia, but unfortunately his assignment of genera into the two groups suggests that he neglected careful examination of these structures for most species. Indeed, genitalia are discussed for only 4 of the 19 peoriine genera included in the paper. The revision treats 30 species in 19 genera of Peoriinae, with 7 of the species and 4 of the genera being described as new.
The great world monograph of Ragonot (1901) was completed by Sir George F. Hampson after the author’s death in 1895 and contains descriptions of 31 species and illustrations in color of 30 species (lotella Hiibner not being illustrated) of North American Anerastiinae placed in 17 genera. Three species and one genus were described as new. A number of the specific names have since been synonymized, and several of the genera have been found not to be North American.
Hampson’s (1918) classification of the subfamily termed the group Hypsotropinae based on Hypsotropa Zeller. He considered dignella Hiibner to be the type of Anerastia, but Ragonot (1901) had already designated lotella Hiibner as such. His paper covers the entire group and ascribes one new species to the North American fauna.
Sixteen new specific names have been added to the literature since the publication of Ragenot’s monograph, these mostly in occasional papers by Dyar (1904, 1904a, 1906, 1908, 1923), Barnes and McDun- nough (1913), Hampson (1918, 1930), and Grossbeck (1917). The most recent checklist for North America (McDunnough, 1939) gives 23 genera and 51 species, plus 1 genus and 10 species as junior synonyms. Heinrich’s ‘‘Revision of the New World Phycitinae’’ (1956) transfers a number of genera across the subfamily lines.
Classification
As has been indicated previously, available information on the Anerastiinae is minimal, so that the present study is only a step toward a more reliable taxonomic system such as exists for many better known groups of insects. Nevertheless, morphological studies of adult specimens indicate that radical changes in the classification of the Anerastiinae are needed.
REVISION OF PEORIINAE AND ANERASTIINAE a
I have accepted the male genitalia as being the most reliable morphological indicators of natural relationships, especially at the generic level. They offer a relatively large number of stable characters. Female genitalia are in many cases too similar to be of much value in separating species, and even genera. Characters of wings, antennae, and palpi previously used to define genera often exhibit more intra- generic than intergeneric variation and consequently are by them- selves extremely unreliable indicators of generic lines.
Numerous examples of wing venation variability could be given, and the following will illustrate the lack of reliability of venational characters: In the genus Peoria the presence or absence in the fore- wing of vein M; does not correlate with any other morphological feature investigated. This vein was present in about one-half of the specimens of P. santaritella, and one specimen possessed M; only on the right wing. Aberrations in the wing venation are very common within the subfamily, and the usefulness of any particular venational feature must be evaluated separately.
Orientation of the palpi may vary with the sex, and in living specimens the position they assume may be different from that found in dead specimens after desiccation. Unfortunately, most species have been described from very small series of specimens, so the variation has not always been apparent. Thus the use of this character in assigning species to genera has proved to be extremely unreliable.
The definition of the Anerastiinae has not generally been regarded as very satisfactory. Although most of the species have a distinctive habitus, this and the reduced tongue are not, in the absence of other substantial correlated characters, sufficient to define the group. Heinrich (1956), following Ragonot’s division of the Phycitidae, felt that ‘. . . in the main the subfamilies [Phycitinae and Anerastiinae] themselves appear to be natural entities, although their definition leaves much to be desired.”
On the basis of the genitalia, the North American Anerastiinae divide into two distinctly separate groups. One of these groups is composed of eight phycitine genera, including Anerastia, which seem to be related to diverse areas of that subfamily, and the remaining seven genera comprise a natural group, the Peoriinae.
The genitalia of both sexes in the peoriines differ distinctly from those in the phycitines. Females all have the ovipositor greatly com- pressed, probably as an adaptation for a particular mode of oviposi- tion. In the males the uncus is so different from its usual appearance in the phycitines as to be nearly unrecognizable in the North American species. Indeed, the assumption that these terminal parts represent uncus is based on an examination of several Old World peoriine
8 U.S. NATIONAL MUSEUM BULLETIN 280
genera in which the uncus is similar, in varying degrees, to that of the phycitines.
A number of other features are generally, but not exclusively, characteristic of the Peoriinae. In all species the males have two to five (possibly more) basal antennal segments fused to some degree, the exact number often varying within a species. Longitudinal wing markings are more common than transverse ones, and the ocelli are well developed in all the species examined.
Inasmuch as the present state of our knowledge is so unsatisfactory, the author hopes to undertake studies of the group on a world basis to better deal with the problem. The interpretation of just how closely the Phycitinae and the Peoriinae are related will depend in part on how homologies between the male genitalic structures of the two groups are interpreted, and in part on the significance of the compressed ovipositor. A better understanding of this problem will require examination of the genitalic structures of all the species, and perhaps studies of their embryology and soft-part anatomy as well. Life history studies would be extremely valuable and should help to elucidate the functional significance of the compressed Ovipositor.
Subfamily Peoriinae ' Hulst Hulst, 1890, p. 102.
Diaenosis.—Similar to Phycitinae; distinguished by spicate modifications of uncus in males and by highly compressed ovipositor of females.
Derscription.—Labial palpi well developed, maxillary palpi mod- erate to very small; male antennae with basal segments of shaft variously fused; tongue reduced; ocelli well developed.
Forewing maculation predominantly longitudinal; transverse bands, when present, usually expressed as dots, more rarely solid; 10 or 11 veins; R, free from cell; R3,, stalked; R; absent; M, free from cell; Cuz free from cell. Hindwing with 6 or 7 veins; Rs closely approximate to or stalked with Sc+R, beyond cell; frenulum of female simple.
Male genitalia with uncus bearing spicate processes. Gnathos various, usually bearing medial process. Transtilla absent (except in Goya). Juxta well developed, various. Vinculum well developed. Eighth abdominal segment without special hair tufts.
Female genitalia with ovipositor strongly compressed. Highth abdominal segment compressed, rarely well expanded anteriorly. Apophyses well developed, approximately equal in length. Ductus
1 Based upon North American species.
REVISION OF PEORIINAE AND ANERASTIINAE 9
bursae moderately short. Bursa well developed, rarely armed. Ductus seminalis from or near posterior end of bursa.
Key to Genera of Peoriinae Based Upon the Male Genitalia
Within the Peoriinae intrageneric variation of alar, palpal, and antennal features seriously limits their usefulness in separating genera. In the absence of other distinguishing features, keys to genera must rely on characters of the male genitalia. Whenever slide preparation is impractical, reasonably certain determinations can often be made by comparing external features of the specimens to descriptions and photographs of adults. Genitalia slides should be prepared if correct identification is critical. It is suggested that the indicated figures be consulted when using the keys.
1. Each spicate process (of uncus) basally branched into two long perpendicular tapering spines of nearly equal length; spicate processes connected by medial
bandlike process (fig. 51)... .. Shed Ase CS Spicate processes unbranched, or one beinch) a least hiwiee iene of other; medial process absent or not bandlike ..... Bian) eet Sem
2. Medial process of uncus bearing prominent SOeteramne cial Peo chnemnees juxta with pair of setaceous tubercles (fig.118) ............ .Anacostia Medial process of uncus unarmed, or with minute cusps only; juxta without pair of setaceous tubercles (figs. 105-117) . ... ; . . . Peoria
3. Spicate processes unbranched, blunt, elbowed in midele: pation tapering to single sharp apical spine (fig. 126) ..... CH 6 os 2 ACASCOSS Spicate processes various, not angled; gnathos mith mie dial process not a single SHATP SPINE 2 < .«. oshe Se alten Meghiatee
4. Each spicate process with Phomboldal fend! pad; ree eilin tical: aedeagus strongly tapered (fig. 131) .... ae ees eae ine OY Rhomboidal pad absent; juxta Seecneuie or shield ahaped: peeets not strongly tapered .... Mie cece ay fae
5. Each spicate process a ered sits Rook Snorer han ae es: arm, tapering to sharp point; gnathos apically tricuspidate (fig. 130). . . . . . Reynosa Spicate processes not as above, at least as long as gnathos arm; gnathos not apically tricuspidate. . ... ee Gest
6. Apical process of gnathos digitate; sais peace eon with (ar lateral and much shorter caudal branches (figs. 119-125) .... joes ont ec REV aCe Gnathos without digitate apical process; spicate peoeeees unbranched (figs. LZ7— EO SIR LIE) 3 Salhi. FE)-Se.. ifn, 4.02) S Homosassa
Peoria Ragonot
Aurora Ragonot, 1887, p. 18; 1889, p. 117. — Hulst, 1890, pp. 209-210. — Smith, 1891, p. 84. — Ragonot, 1901, p. 337. — Hulst, 1902, p. 437. — Barnes and McDunnough, 1917, p. 149.— Hampson, 1918, p. 106.— Forbes, 1923, p. 638.— McDunnough, 1939, p. 35. [New synonymy. Type: Aurora longipalpella Ragonot, 1887. Monobasic.]
Peoria Ragonot, 1887, p. 19; 1889, p. 117. — Hulst, 1890, p. 213. — Smith, 1891, p. 85. — Ragonot, 1901, p. 386. — Hulst, 1902, p. 439. — Barnes and
10 U.S. NATIONAL MUSEUM BULLETIN 280
McDunnough, 1917, p. 149. — Forbes, 1923, p. 638. — McDunnough, 1939, p. 36. [Type: Anerastia haematica Zeller, 1872. Original designation.]
Statina Ragonot, 1887, p. 19; 1889, p. 117. — Hulst, 1890, p. 216. — Smith, 1891, p. 85. — Ragonot, 1901, p. 415. — Hulst, 1902, p. 440. — Barnes and McDunnough, 1917, p. 150. — Hampson, 1918, p. 59. — McDunnovugh, 1939, p. 36. [New synonymy. Type: Statina roseotinctella Ragonot, 1887, p. 19. Monobasic.]
Calera Ragonot, 1888, p. 50.— Smith, 1891, p. 85. — Ragonot, 1901, p. 417. — Hulst, 1890, p. 217. — Barnes and McDunnough, 1917, p. 150. — Hamp- son, 1918, p. 59. — Hulst, 1902, p. 441. — Forbes, 1923, p. 639. — McDun- nough, 1939, p.36. [Newsynonymy. Type: Calera punctilimbella Ragonot, 1888, p. 50. Monobasic.]
Altoona Hulst, 1888, p. 116; 1890, pp. 206-207. — Smith, 1891, p. 84. — Hulst, 1902, p. 488. [New synonymy. Type: Anerastia opacella Hulst, 1887. Original designation.]
Cayuga Hulst, 1888, p. 116; 1890, pp. 208-209. — Smith, 1891, p. 84. — Hulst, 1902, p. 438. [New synonymy. Type: Spermatophthora gemmatella Hulst, 1887. Monobasic.]
Volusia Hulst, 1890, p. 206 [not Robineau-Desvoidy, 1830, p. 674, in Diptera, Ortalidae; not Adams, 1861, p. 306, in Gastropoda]. — Smith, 1891, p. 84. [New synonymy. Type: Volusia roseopennella Hulst, 1890. Monobasic.]
Wekiva Hulst, 1890, p. 215. — Smith, 1891, p. 85. — Hulst, 1902, p. 440. [New synonymy. Type: Wekiva nodosella Hulst, 1890. Monobasic.]
Osceola Hulst, in Smith, 1891, p. 85 [not Baird and Girard, 1853, p. 133 p. 183, in Reptilia]. [Nomen nudum. New synonymy. Type: Chipeta perlepidella Hulst, 1892. Monobasic.]
Chipeta Hulst, 1892, p. 62. — Barnes and McDunnough, 1917, p. 150. — McDun- nough, 1939, p. 36. [New synonymy. Type: Chipeta perlepidella Hulst. 1892. Monobasic.]
Chipota [sic] — Hulst, 1902, p. 441. [Misspelling for Chzpeta.]
Trivolusia Dyar, 1902 [1903], p. 438. [Newsynonymy. Type: Volusia roseopen- nella Hulst, 1890. Monobasic.]}
Ollia Dyar, 1904, pp. 107-108. — Barnes and McDunnough, 1917, p. 149. — Mc- Dunnough, 1939, p. 36. [New synonymy. ‘Type: Ollia santaritella Dyar, 1904. Monobasic.]
Dracnosis.—Members of this rather variable genus may be recog- nized by the characteristic uncus with its bandlike medial process and paired spicate processes, bifurcate with each arm round, slender, sharply pointed, and oriented at about 90 degrees to the other (fig. 51).
Derscription.—Labial palpi porrect (obliquely ascending in lutei- costella); tongue rudimentary; antennae with base compressed, male shaft with basal segments partly fused, female with shaft filiform, finely ciliate.
Forewings with 10 or 11 veins; R, from cell well before upper outer angle; R» from cell near base of or stalked with R344; Ray, always stalked, from just before the angle; M, from the angle; M; stalked or fused with M;; M; sometimes stalked with Cu,; Cu, at or near lower outer angle of cell; Cuz from just before the angle. Hindwings with 6 or 7 veins; Se and Rs closely approximate or stalked; M, from upper
REVISION OF PEORIINAE AND ANERASTIINAE 11
outer angle of cell; Mz absent; M; stalked with Cu, for at least one- third of its length, sometimes fused; Cu, from lower outer angle; Cuz from just before the angle.
Male genitalia with uncus bearing bandlike medial process; spicate processes each terminating in slender, sharply pointed ventrad and caudad arms. Gnathos arms broad and flat. Vinculum well developed, broadly rounded. Aedeagus somewhat flattened, three to four times as long as broad; vesica, when armed, bearing one or two cornuti.
Female genitalia with posterior and anterior apophyses of about equal length, well developed. Ductus bursae moderately short. Bursa unarmed. Ductus seminalis from posterior end of bursa, broadened at base.
Discussion.—The genus includes 13 known species and is the largest in the American fauna of the subfamily. Affinities vary within the genus, and the species have been grouped to show relationships in so far as this is possible with a linear arrangement.
There is enough variation of characters within many of the species to render difficult the construction and use of keys. To aid in the identi- fication of specimens, a chart summarizing the characters found to be of greatest value in separating species is included in addition to the key.
In a few cases the male genitalia are quite distinctive of the species, but for the most part the differences are rather subtle. The form of the gnathos, number of cornuti in the vesica, and shape of the valvae (not summarized in table, see figures) are the most useful characters of the male genitalia.
Navasota hebetella may belong to Peoria and has been included in the key (see Unplaced Genera and Species, p. 87).
In accordance with the recommendation of the Code (Article 24), I have chosen the name Peoria for this genus rather than the lesser known Aurora which has page precedence.
Explanation of Table 1
1. Forewing, vein Re x=free from cell o=stalked with R314 2. Forewing, vein M; x=stalked with M3; *=fused with M; (apparently absent) 3. Forewing, vein Cu; x= free from cell o=stalked with M; 4. Forewing, region anterior to cell x=abruptly pale, with few or no darker scales o=not abruptly pale, Sc and Rs sometimes white traced
285—-934—68——2
12
a
10.
Hots
12.
U.S. NATIONAL MUSEUM BULLETIN 280
Forewing x= ground darker posterior to cell, at least at base o= ground not darker posterior to cell Forewing x=some or all veins white traced o= without white tracing on veins Forewing, transverse posterior line
x= present *— represented by dots o= absent Labial palpi p=porrect
a= ascending Maxillary palpi x=reaching frons or nearly so o=small, not approaching frons Male antennae 1=laminate S=serrate ss=subserrate Gnathos of male genitalia a= with apical process l= with pair of subapical processes o= without special processes Aedeagus, number of cornuti in vesica
The use of two symbols separated by a comma indicates two characters appear- ing with about equal frequency. A symbol in parentheses indicates a character appearing less frequently than its alternative.
TABLE 1.—Summary of characters found to be the most useful in distinguishing species of Peoria*
Species 1 2 3. OA 5 6 1 8°°9 longipalpella oO x xO oO oO <1 oD bipartitella O * 8) C9) S@)- S@) OV 10 tetradella xe OX me Sx oO x ORD opacella x OS xa) x x o p floridella x50) 0X xq0O X XG @) 4 rostrella oO x x {10 oO x OW Dp gemmatella x x x0 oO X o p roseotinctella oO a Oo x,0 oO x xp johnstont oO s xi) oO x 01) Dp santaritella x foxy | x ao x oO OmD holoponerella x Xi xe oO oO o p approximella oO * Kauex oO oO OM Dp luteicostella oO * KEK oO oO Oo a
| |
*See explanation on preceding pages.
OM (O40! 16) 6) 16) IH! (OPA) Id ivi
10
ss
5S
12
KSB OrR KF NNN KR RRR Re
bo
10.
ine:
12.
13.
14,
REVISION OF PEORIINAE AND ANERASTIINAE 13
Key to the Species of Peoria Based Upon External Features
R, stalked with R344; Mz absent . A
R, not stalked with Rs+4; if stalked, fen M 2 reece a M; stalked with Cu;; hindwing M; absent Ors; M; not stalked with Cu, sometimes from point; ae M; erie
ww bo
DRCSCIUs at el ar? hh evahe' prayvayseagairy ok 4 Transverse poeenor udicated ne ine of dots Ga 10). sn ei Se oneeupuetelln Transverse posterior absent; terminal line of dots present . .N. hebetella Forewing abruptly pale anterior to cell, with no more than a few scattered
nonwhite scale. ... . FPS ASP PHIL NL CBS IONG Forewing not abruptly pale eats coil ee OF. CORE Riss. 3 eG Forewing pale posterior to A2; palpi porrect (fig. 14) .. . . .approximella Forewing not pale posterior to A2; palpi ascending (fig. 12) . . luteicostella Forewing darker posterior to cell; mn es palpi approaching frons; (fig.
Seer oS a es eee Dipartivella Forewing not darker Beceem e cell; Vansitines palpi not approaching
EONS Gigs 1) oo ses AD Lag eee sa ohustont Forewing with transverse paode (Gg. 9). see gh tep cb o¥ies olongipalpella Transverse bands absent .. . oe abategs Soke Forewing light buff, abruptly pale aeane 3 cell ey 3. i . holoponerella Forewing not abruptly pale anterior to cell . .........2.2.-49 Weims traced’ with white; ¢ 0. « + te ae et, ey SOE Veins not traced with white. ... . eer tea rat |!) Ground white anterior to cubitus; eter peered Feels with red;
maxillary palpi not reaching frons (fig. 5) . 2) Suse c-e aust ¢o) Santaritella
Ground a mixture of brown and white eles: aeeliy lighter anterior to cubitus, at least near base; aio palpi reaching frons (fig.
1M) Rey Sue: sea a ho 3 Se Opacella Forewing ne eee ones to oa nb Teaet near poe Set oh ALAYB etek Forewing not darker posterior to cell. . .. . esas eden NS A, traced with white, bordered with brown; eae Pain not reaching frons) (figs.8)) 5 =. . . . . floridella
Ag not traced with white: Prcillenry ta peace eon (Ge Pt); . opacella Cubitus and lower outer angle of cell broadly traced with white (fig.
6) . wh Os Parcy eee tay Mra dente ... . . tetradella Not as poere: Se na Rs traced with: cee ronal aoa PEO WIL oe, <<) oor naulk R, free from cell (fig. 7). eee hie ee Oe ah ae cae ee SORIMBLCLA Ry stalked with Ra;+« (fig. 2) Bra Mett ey at get cn Star fla or epee cook OSTECLG:
Peoria longipalpella (Ragonot), new combination Fiaurss 9, 51, 70, 105, 144
Aurora longipalpella Ragonot, 1887, p. 18. — Hulst, 1890, p. 210. — Smith, 1891,
p. 84. — Ragonot, 1901, pp. 337-338. — Hulst, 1902, p. 437. — Barnes and McDunnough, 1917, p. 149. — Hampson, 1918, p. 106. — Forbes, 1923, p, 638. — McDunnough, 1939, p. 35. — Kimball, 1965, p. 250.
Diacnosis.—This is the only known species of Peoria in which the
forewings are marked with unbroken transverse bands, although transverse rows of dots occur in a few other members of the genus.
14 U.S. NATIONAL MUSEUM BULLETIN 280
Description.—Frons brown; labial palpi with basal segments white, second and third segments brown on outer sides, white ventrally and on inner sides; maxillary palpi reaching frons, brown; antennae brown, male subserrate; occiput laterally, patagia, and tegulae brown, vertex and occiput light brown dorsally.
Forewings with ground white anterior to cubitus, brown posterior to cell; cell sprinkled with brown, with orange red anterior to cell; transverse anterior and posterior bands brown; ground brown on underside, bands darker; with 11 veins; Re stalked with Rgi4; Mo43 stalked. Hindwings light brown; 7 veins; Sc and Rs stalked; M3, stalked with Cu.
Male genitalia with gnathos bearing apical digitate projection. Juxta U-shaped. Valvae with costa sparsely setose, terminating in sharp projecting tooth; sacculus densely pubescent. Aedeagus with vesica bearing a single serrate cornutus.
Female genitalia with ovipositor tip moderately setose, caudal margin with numerous fine setae, sparsely setose laterally. Apophyses curved at base, posterior with base pointed and slightly flattened; anterior with base thickened.
Tyrr.—In the British Museum (Natural History).
TypE paTa.—Lectotype female, hereby designated, ‘667; U.S.A.; n. gen. R. longipalpella Rag’’; genitalia slide No. 707, J. Shaffer, Jan. 4, 1967. The specimen is glued together between the meso- and metathorax. Type data given with the original description consists of: “QO 22 mill.” Ragonot (1901) in a presumed reference to the type reports: ‘“Amerique boreale, 11 octobre. Une 9. Mus brit.”
SPECIMENS EXAMINED.—2 0", 2 9.
DistripuTIoN (Map 8).—Recorded from North Carolina and Virginia.
UNITED STATES: Norrs Carouina, Polk Co., Tryon, 1 ?, no date (Fiske), [USNM]; Wake Co., Raleigh, 1 9, ““M. Aug. 09,” [USNM].
Virainia: Arlington Co., Arlington, 1 o&, Aug. 28, 1951 (J. G. Franclemont),
[USNM]. Locality UNKNOWN: 1, no date, Sweadner collection [CM].
Discussion.—Kimball reports the species from Siesta Key, Fla., May 13, 1946, but I have not seen the specimen.
Peoria bipartitella Ragonot Ficures 13, 71, 106, 145
Peoria bipartitella Ragonot, 1887, p. 19; 1889, p. 117. — Hulst, 1890, p. 214. — Smith, 1891, p. 85. — Ragonot, 1901, p. 386. — Hulst, 1902, p. 489. — Barnes and McDunnough, 1917, p. 149. — McDunnough, 1939, p. 36. — Kimball, 1965, p. 251.
REVISION OF PEORIINAE AND ANERASTIINAE 15
Volusia roseopennella Hulst, 1890, p. 206. — Smith, 1891, p. 84. — Rindge, 1955, p. 170. [New synonymy.]
Tolima roseopennella (Hulst). — Ragonot, 1901, p. 340. — Barnes and MeDun- nough, 1917, p. 149. — McDunnough, 1939, p. 35. — Kimball, 1965, p. 250.
Trivolusia roseopennella Hulst, 1902, p. 438. — Grossbeck, 1917, p. 134.
Commotria roseopennella (Hulst). — Hampson, 1918, p. 108.
Hypsotropa bipartitella (Ragonot). — Hampson, 1918, p. 78.
Peoria bipunctella Ragonot. — Forbes, 1923, p. 639. [Not Ragonot.]
Dracnosis.—The presence of a well-developed digitate medial process on the gnathos serves to delimit this species from among those of the genus with similar wing venation.
DescripTion.—Frons conical, varying brown to light pink; labial palpi with basal segments white, outer sides of second and third segments varying brown to light pink, inner sides with white-tipped scales; maxillary palpi approaching frons, brownish white; antennae light brown, laminate in male; occiput brown, vertex white behind antennae, patagia and tegulae varying grayish red to brown, legs grayish red to brown on outer sides, white on inner sides.
Forewings with extremely variable maculation; area anterior to cell red, brown, white, or mixtures of scales of these colors; cell usually red or white, brown in some (Mississippi) specimens, often a mixture of these three colors; area posterior to cell usually red, sometimes brown; veins traced with white, tracing variable, sometimes absent; 10 veins, Re stalked with R3,4; Me,3 fused, from lower outer angle. Hindwings light brown, darker in apical area; 7 veins; M; and Cu, long stalked.
Male genitalia with gnathos bearing apical digitate process. Juxta V-shaped. Vinculum with anterior margin flat and flanged. Valvae with costa sparsely setose, bearing blunt apical tooth; sacculus with fine hairs. Aedeagus with vesica bearing single subserrate cornutus.
Female genitalia with ovipositor finely setose on caudal margin, moderately setose laterally. Apophyses curved, posterior curving upward at base and tapering to point, anterior slightly thickened at base.
Typres.—P. bipartitella, in the Museum National d’Histoire Naturelle; V. roseopennella, in the American Museum of Natural History.
Type pata.—P. bipartitella, lectotype female, hereby designated, North Carolina, Morrison; genitalia slide No. 534, J. Shaffer, May 14, 1965; V. roseopennella, lectotype male, hereby designated, Florida, March; genitalia slide No. 3220, Carl Heinrich, June 14, 1946; the original description reports “Volusia County, Fla.,’’ no date.
REcorDED HostT.—Poaceae: Panicum lanuginosum Ell. from slide label in USNM, specimens taken July 8, 1950, Kill Devil Hills, N.C.
SPECIMENS EXAMINED.—34 o’, 19 9.
16 U.S. NATIONAL MUSEUM BULLETIN 280
Distripution (Map 8).—Chiefly the Atlantic and Gulf Coastal Plain, recorded from Mississippi to Massachusetts, also from north central Illinois.
Fernald (1950) reports the food plant as occurring in ‘‘Sandy open soil, thin woods, etc., Fla. to Tex., n., rather locally, to s. R.I.”
UNITED STATES: Ftoripa: Alachua Co., Gainesville, 2¢°, July 8, 1927 (J. Speed Rogers) [CU, USNM]; 17, July 1927 (J. Speed Rogers) [CU]; High- lands Co., Lake Placid, 19, Mar. 6, 1945 (J. G. Needham) [CU]; Manatee Co., Myakka City, 19, Feb. 17, 1945 (J. G. Needham) [CU]; Sarasota Co., Siesta Key, 107, Feb. 28, 1951 (C. P. Kimball) [CNC].
Intrnois: Putnam Co., 19, Aug. 29, 1961 (M. O. Glenn) [MOG]; 19, Aug. 1, 1964;19, Aug. 29, 1964.
MassacuvuseEtts: Barnstable Co., Barnstable, 1 <7, July 10, 1949 (C. P. Kimball) [CNC]; 14,29, July 11, 1949 (C. P. Kimball) [CNC, CPK]; 2 2, July 14, 1949 (C. P. Kimball) [CNC]; 1¢, July 15, 1949 (C. P. Kimball) [CPK]; West Barn- stable, 2 o, 19, July 16, 1949 (C. P. Kimball) [CNC, CPK]; Barnstable, 107, July 11, 1950 (C. P. Kimball) [CPK]; 1<7, July 28, 1950; 19, Aug. 7, 1952; 1, July 4, 1958; Dukes Co., Martha’s Vineyard, 1?, Aug. 1, 1926 (F. M. Jones) [CNC]; 10, Aug. 8, 1941 (F. M. Jones) [USNM]; 19, Aug. 4, 1944 (F. M. Jones) [ANS]; 17, Aug. 6, 1944; 10, Aug. 6, 1946 (F. M. Jones) [CNC]; 107, Aug. 4 (F. M. Jones) [USNM]; 19, Aug. 21 (F. M. Jones) [USNM]; Vineyard Haven, 1, July 17 (F. M. Jones) [ANS]; Plymouth Co., Agric. Exp. Sta., East Wareham, 13, July 11, 1962 [CPK].
Mississippi: Forrest Co., Camp Shelby, 6 o, 3 9, Sept. 1-15, 1944 (C. D. Michener) [AMNH]; 2 &, Sept. 16-30, 1944.
New Jersey: Bergen Co., Oakland, 1 0, Aug. 5, 1948 (C. P. Kimball) [CPK]; 1 o&, Aug. 13, 1948 (C. P. Kimball) [CNC]; 1 9, Aug. 14, 1948; Burlington Co., Whitesbog, 1 9, July 12, 1939 (KE. P. Darlington) [ANS]: 1 o, Aug. 2, 1940.
NortaH Carouina: Macon Co., Horse Cove, Highlands, 1 o&, Aug. 3, 1957 (C. J. Curen) [CNC]; Polk Co., Tryon, 1 &, Aug. 18, 1903 (Fiske) [USNM]; 1 9, no date.
VirGiInta: Nasemond Co., Holland, 1 9, Aug. 11, 1944 (O. Buchholz) [ANS].
Peoria tetradella (Zeller), new combination Ficurss 6, 72, 107, 146
Anerastia tetradella Zeller, 1872, pp. 552-553.
Saluria tetradella (Zeller). — Ragonot, 1889, p. 117; 1901, pp. 362-363. — Hulst, 1902, p. 439. — Barnes and McDunnough, 1917, p. 149. — Hampson, 1918, p. 102. — MeDunnough, 1939, p. 36.
Altoona tetradella (Zeller). — Hulst, 1890, p. 207. — Smith, 1891, p. 84.
Draenosts.—The broad white band on the cubitus and lower outer angle of the cell marks this species as distinct from other members of the genus.
Derscription.—F rons brown; labial palpi with basal segments white, second and third segments gray on outer sides, white on inner sides and ventrally; maxillary palpi reaching frons, light brown; antennae light brown, male subserrate; occiput laterally, patagia, and tegulae
REVISION OF PEORIINAE AND ANERASTIINAE 17
light brown, occiput dorsally and vertex white, legs white on inner sides, outer sides clothed with white-tipped brown scales.
Forewings with white band anterior to cell tapering to point at apex, sprinkled with reddish-brown scales, bordered posteriorly by dark brown band of varying width, often extending to A, fold, fre- quently scattered with reddish-brown scales; reddish brown posterior to A, fold, A; white traced and bordered with dark brown scales; veins traced with white, very prominently so at lower outer angle of cell; 11 veins; R» from cell close to base of or stalked with R34; Mo.3 stalked. Hindwings light brown, darker toward apex; with 7 veins.
Male genitalia with gnathos bearing apical digitate projection. Juxta V-shaped. Vinculum flanged and thickened at apex. Valvae pointed apically; costa sparsely setose; sacculus with fine hairs. Aedea- gus with vesica bearing single serrate cornutus.
Female genitalia with ovipositor bearing numerous fine setae on caudal margin, tip moderately setose. Apophyses curved, posterior tapering to point.
Typr.—lIn the British Museum (Natural History) (from the Zeller collection).
Typr pata.—In the original description Zeller notes: ‘‘Vaterland: Texas (Boll, Belfrage). Mehrere o& und @ fing Belfrage in der ersten Hialfte des June, 2 Q am 8. und 9. Juli.”
Lectotype male, hereby designated, labeled: ‘10/6, Bosque Co Texas, Tetradella Z. Texas [green label in Zeller’s handwriting], Zell. Coll. 1884, o& genitalia slide 6-II-1967 J. Shaffer No. 728.”
Lectoparatypes: One male and four females, all labeled Bosque County, Tex. and bearing Zeller’s personal type label; male dated 11/6 (June 11), two females dated 7/6, other two females dated 8/7 and 9/7.
RECORDED HosT.—Poaceae: Elymus canadensis L., borer in stem. From specimen label, three males, Lafayette, Ind.
The host occurs throughout most of the United States and southern Canada. Hitchcock (1951) notes: “River banks, open ground, and sandy soil, Quebec to southern Alaska, south to North Carolina, Missouri, Texas, Arizona, and northern California.”
SPECIMENS EXAMINED.—22 go’, 41 Q.
DistrisuTion (Map 6).—California to Texas, northeastward to Ontario.
UNITED STATES: Catirornia: Modoc Co., Canby, 2 o, July 20, 1927 [CU]; county unknown, ‘‘Mad R. ab. Maple Cr.,” 1 o&, July 28-29, 1927 [CU].
ILuino1s: Putnam Co., 1 9, Aug. 6, 1958 (M. O. Glenn) [MOG]; 1 o&, June 21, 1961; 1 o, July 10, 1961; 1 9, July 12, 1961;1 9, July 17, 1961;1 9, June 24, 1962; 1 9, July 5, 1962; 1 @, June 23, 1963; 1 9, July 2, 1964.
INDIANA: Tippecanoe Co., Lafayette, 3 o@, 1916 (C. N. Ainslie) [USNM].
Kansas: Thomas Co., (3150 ft.), 1 o&, no date (F. X. Williams) [UK].
18 U.S. NATIONAL MUSEUM BULLETIN 280
Texas: Blanco Co., 1 9, May [USNM];1 9, June [AMNH];1 0,1 9, July [AMNH]; 1 9, August [AMNH]; 2 9, no date, [USNM]; Bosque Co., 1 o, “11/6” (Zeller collection) [BM]; Burnet Co., 2 o&@, no date (F. G. Schaupp) [USNM]; Collin Co., Plano, 4 9, July (E. 8. Tucker) [USNM]; Dallas Co.,1 9, June [AMNH]; Kerr Co., Kerrville, 1 9, May 30, 1906 (F. C. Pratt) [USNM]; 3 9, May 31, 1906; 1 o, 2 9, June 1, 1906; 1 9, Apr. 11, 1907; 1 9, no date; La Salle Co., Cotulla, 1 9, May 12, 1906 (Crawford and Pratt) [USNM]; Medina Co., Sabinal River opposite Hondo, 1 &@, July 1, 1917 [CU]; county unknown, “Black Jack Spgs.,” 1 9, no date [USNM]; locality unknown, 5 ? [USNM]; 3 0,5 9 [INHS].
CANADA: Onrarto: Port Colborne, 1 @, June 24, 1934 (J. J. de Gryse) [CNC].
Peoria opacella (Hulst), new combination Fiaures 11, 73, 108, 147 Anerastia opacella Hulst, 1887, p. 138. — Rindge, 1955, p. 168. Altoona opacella Hulst, 1887, p. 116; 1890, p. 207. — Smith, 1891, p. 84. — Hulst, 1902, p. 438. Saluria dichroeella Ragonot, 1889, pp. 118, 117. — Barnes and McDunnough, 1917, p. 149. — McDunnough, 1939, p. 36. [New synonymy.] Tolima opacella (Hulst). — Ragonot, 1889, p. 117; 1901, p. 341. — Barnes and McDunnough, 1917, p. 149. — McDunnough, 1939, p. 35. Altoona dichroeella (Ragonot). — Hulst, 1890, p. 207. Altoona dichrocella [sic]. — Smith, 1891, p. 84. Saluria dichrocella Hampson in Ragonot, 1901, p. 363; 1918, p. 101. [New synonymy, objective.] Saluria dichroella [sic]. — Hulst, 1902, p. 439. Commotria opacella (Hulst). — Hampson, 1918, p. 108.
Draanosis.—This species is distinguished from all other members of the genus with similar venation by the combination of the forewing being basally darker posterior to the cell and the absence of white tracing on forewing vein A».
Description.—Frons oblique, scales brown with white tips; labial palpi with basal segments white, second and third segments clothed with white-tipped brown scales dorsally and on outer sides, with light brown to white scales ventrally and on inner sides; maxillary palpi almost reaching frons; antennae brown, laminate in male; occiput behind eye, patagia, and tegulae brown, vertex behind antennae and dorsum of occiput light brown; legs with tarsi clothed with white- tipped brown scales.
Forewings brown, darker posterior to cubitus, especially basal half; all veins except anals variously traced with white; areas between veins a mixture of white and brown scales; usually solid brown poste- rior to cubitus, becoming lighter distally. Hindwings light brown, slightly darker in apical region. Venation as in tetradella.
Male genitalia with gnathos bearing apical digitate projection. Juxta V-shaped. Vinculum with anterior edge flattened and flanged. Valvae with costa sparsely setose, sometimes with short apical tooth;
REVISION OF PEORITINAE AND ANERASTIINAE 19
sacculus with numerous fine hairs. Aedeagus with vesica bearing a single serrate cornutus.
Female genitalia with ovipositor tip and caudal margin moderately setose, sparsely setose laterally. Posterior apophyses curved on basal third, base flattened, tear-shaped, tapering to point; anterior slightly curved.
Typrs.—A. opacella, in the American Museum of Natural History (lectotype), in the U.S. National Museum (lectoparatype); S. di- chroeella, in the Muséum National d’Histoire Naturelle.
Type pata.—A. opacella, lectotype male, hereby designated, Blanco Co., central Texas, collection G. D. Hulst, genitalia slide No. 3221, Carl Heinrich, June 14, 1946.
Lectoparatype: Texas, one female, Fernald collection; USNM 40080; genitalia slide No. 1107, Carl Heinrich, Apr. 5, 1938.
S. dichroeella, lectotype male, hereby designated, Texas, October, genitalia slide No. 533, J. Shaffer, May 14, 1965.
SPECIMENS EXAMINED.—47 <’, 46 Q.
DistRIBUTION (Map 7).—Arizona, New Mexico, southwestern Texas to Brownsville.
UNITED STATES: Arizona: Cochise Co., Cochise Stronghold,1 9? , Aug. 30, 1958 (P. Opler) [UCB]; Ramsay Canyon, Huachuca Mts., 1 9, July 10-15, 1941 (A. B. Klots) [AMNH]; Southwestern Res. Sta., 5 mi. west Portal (5400 ft.), 1 o, July 7, 1956 (Cazier and Ordway) [AMNH];1 9, July 9, 1956; 1 9, July 20, 1957 (M. Statham) [AMNH]; 1 0, Aug. 2, 1956 (C. and M. Cazier) [AMNH]; 1 o&, July 26-Aug. 3, 1959 (A. B. Klots) [ABK]; Cococino Co., 64 mi. EESE. Flagstaff (6500 ft.), 1 @, Aug. 7, 1964 (J. G. Franclemont) [JGF]; Vail Lake Rd., 9% mi. SE. Flagstaff (6500 ft.), 1 9, July 11, 1961 (J. G. Franclemont) [JGF]; 1 9, July 18, 1961 (R. W. Hodges) [JGF]; Pima Co., Baboquivari Mts., 4 9, July 1-15, 1923 (O. C. Poling) [USNM]; 13 o&, 4 9, July 15-30, 1923; 1 9, July 27-31, 1923; 3 9, Aug. 1-4, 1923; 7 9, Aug. 1-15, 1923; 3 9, Aug. 15-30, 1923; 2 9, Aug. 24-31, 1923; 1 9, Sept. 1-15, 1923; 2 9, July 1-15, 1924; 5 9, Aug. 15-30, 1924; 1 9, Sept. 15-30, 1924; 2 9, no date, (F. H. Snow) [UK]; 1 ?, no date (F. H. Snow) [USNM]; Pinal Co., Desert Arboretum, Superior, 1 o, July 18-21, 1941 (A. B. Klots) [AMNH}]; Santa Cruz Co., Santa Rita Mts., Madera Canyon (4880 ft.), 1 @, July 9, 1959 (J. G. Franclemont) [JGF]; 10, Aug. 23, 1959.
New Mexico: Eddy Co., White City, 2 #, May 15, 1950 (E. C. Johnston) [CNC]; 1 o&, May 16, 1950; 1 @, May 17, 1950; 1 ?, July 23, 1959 (A. B. Klots) [ABK].
Texas: Brewster Co., Alpine, 2 &', May 22, 1950 (E. C. Johnston) [CNC]; Cameron Co., Brownsville, 1c, no date [USNM]; San Benito, 407, March 16-23 [USNM]; 16, March 24-30; 14, 19, July 16-23; 19, July 24-31; 1 #, nodate; Hidalgo Co., Mercedes, 2 #, Aug. 31, 1958 (H. Smalzried) [AMNH]; Jeff Davis Co., Ft. Davis, 27, May 20, 1950 (E. C. Johnston) [CNC]; Limpia Canyon, 8 7 1 9, May 20, 1950 (E. C. Johnston) [CNC]; 1 &, June 4, 1950.
Discusston.—The species shows close affinities to tetradella and floridella, particularly to the latter.
20 U.S. NATIONAL MUSEUM BULLETIN 280
Wing maculation in opacella is rather variable, especially with respect to the degree of white tracing on the veins. The variation is apparent within local populations and does not appear to be geographic.
Peoria floridella, new species Figures 8, 55, 74, 109
Draenosts.—The species is rather similar to opacella in most re- spects, but is easily distinguished by the white trace on forewing vein Ao.
Derscription.—Frons conical, scales solid light brown; labial palpi with basal segments white, second segments reddish brown on outer sides, third segments brown on outer sides, both white on inner sides; maxillary palpi moderately small, not approaching frons; antennae light brown, male laminate; occiput behind eye, patagia, and tegulae light brown, vertex behind antennae brownish white; legs white, tarsi light brown.
Forewings reddish orange anterior to cell; ground terra cotta; costa and all other veins traced with white, trace of A, bordered with dark brown, cubitus bordered posteriorly with dark brown line. Hindwings light brown, darker toward apex. Venation as in fetradella.
Male genitalia with gnathos bearing apical digitate projection. Juxta V-shaped. Valvae with costa sparsely setose, bearing poorly de- veloped apical tooth; sacculus with numerous fine hairs. Aedeagus with vesica bearing a single serrate cornutus.
Female unknown.
Types.—In the Canadian National collection (holotype); in the American Museum of Natural History (one paratype), in the Carnegie Museum (two paratypes), in the collection of Charles P. Kimball (one paratype).
Type pata.—Holotype, male, Volusia County, Fla., Aug. 2, 1956, H. A. Denmark; C.N.C. Type No. 9439; genitalia slide No. 412, J. Shaffer, Mar. 15, 1965.
Paratypes: One male, Summer Haven, St. Johns County, Fla., May 30, 1950 (Fred H. Rindge) [AMNH]; one male, Pellicer Cr., 13 mi. north of Bunnell, Flagler County, Fla., Apr. 11, 1954 (J. Bauer), Carnegie Museum Acc. 17023; one male, Apr. 22, 1954, genitalia slide No. 521, J. Shaffer, May 2, 1965; one male, same data as holotype, abdomen lost [CPK].
OTHER SPECIMENS EXAMINED.—15 ©.
DistTrIBuTION (Map 10).—Known only from the east coast of Florida.
UNITED STATES: Fiorina: Indian River Co., Vero Beach, 13 6, April 1941 (J. R. Malloch), USNM; 1 o, May 1941; 1 o, Nov. 15-Dee. 31, 1941.
Discusston.—This form has its closest affinities to opacella, and differs mainly in size and in that the valvae are rounded and lack the
REVISION OF PEORIINAE AND ANERASTIINAE Ft
pointed apex of that species. The most obvious distinctions are those of size, floridella having an alar expanse of 25 mm to 28 mm in the specimens examined, contrasted with about 18 mm to 22 mm for opacella, and maculation, the forewing veins all showing a prominent white trace in the former species. In opacella the white trace is less distinct and essentially absent on Az. The maxillary palpi are moder- ately short in the Florida species, not nearly attaining the frons as in opacella, and the frons itself is clothed with white-tipped brown scales in the latter species rather than the solid brown scales of floridella.
Peoria rostrella (Ragonot), new combination Ficures 2, 75, 110, 148
Saluria rostrella Ragonot, 1887, p. 18; 1889, p. 117. — Hulst, 1890, p. 211. — Smith, 1891, p. 85. — Ragonot, 1901, p. 363. — Hulst, 1902, p. 439. — Barnes and McDunnough, 1917, p. 149. — McDunnough, 1939, p. 36.
Draenosis.—The combination of reddish-brown forewings, veins traced with white, R»2 well stalked with R3.,, and Mz well stalked with M; identify this from within the genus.
Description.—Frons conical, reddish brown; labial palpi with basal segments white, second and third segments brown on outer sides, light brown on inner sides; maxillary palpi cylindrical, reaching frons, light brown; antennae light reddish brown, subserrate in male, cilia about one-half as long as segment width; occiput, patagia, and tegulae reddish brown, vertex somewhat lighter.
Forewings reddish brown; all veins traced with white, A; traced on distal third; 11 veins; Re well stalked with R34; Me well stalked with M3. Hindwings with 7 veins; Sc and Rs stalked; M; stalked with Cu, for about one-half its length.
Male genitalia with gnathos bearing well-developed medial digitate process. Juxta U-shaped. Valvae with costa unarmed. Aedeagus with vesica bearing a single small serrate cornutus.
Typr.—In the Muséum National d’Histoire Naturelle.
Type pata.—Lectotype male, hereby designated, genitalia slide No. 552, J. Shaffer, July 28, 1965. The type locality is given in the original description as California.
OTHER SPECIMENS EXAMINED.—4 o, 1 Q.
Distrisution (Map 6).—Washington southward to central California.
UNITED STATES: Cauirornia: Sonoma Co., Forestville, 1 @, July 10, 1935 (BK. C. J.) [CNC]; Tuolumne Co., Twain Harte, 1 9, Aug. 18, 1960 (M. Lundgren) [UCB]; 1 ¢, July 20, 1961.
WASHINGTON: Grant Co., Dry Falls, 1 o, June 30, 1949 (E. C. Johnston) [CNC]; county unknown, Berne, 1 &@, July 12, 1942 (EK. C. Johnston) [CNC].
22 U.S. NATIONAL MUSEUM BULLETIN 280
Discussion.—The forewing color and maculation is very similar to that of P. johnston.
Peoria gemmatella (Hulst), new combination Fiaurss 7, 76, 111, 149
Spermatophthora gemmatella Hulst, 1887, p. 134. — Rindge, 1955, p. 163.
Cayugag emmatella Hulst, 1888, p. 116; 1890, p. 209. — Smith, 1891, p. 84.—Hulst, 1902, p. 438.
Poujadia gemmatella (Hulst).— Ragonot, 1889, p. 117.
Cayuga bistriatella Hulst, 1890, p. 209. — Smith, 1891, p. 84. — Rindge, 1955, p. 158. [New synonymy.]
Pectinigeria gemmatella (Hulst).— Ragonot, 1901, pp. 356-357. — Barnes and Mc- Dunnough, 1917, p. 149. — Forbes, 1923, p. 638. — McDunnough, 1939, p. 35.
Pectinigera [sic] bistriatella (Hulst). — Dyar, 1908, pp. 117-118.
Pectinigera [sic] pamponerella Dyar, 1908, p. 117. [New synonymy.]
Pectinigeria bistriatella (Hulst).—Barnes and MeDunnough, 1917, p. 149.—Mce- Dunnough, 1939, p. 35. — Kimball, 1965, p. 250.
Pectinigeria pamponerella Barnes and McDunnough, 1917, p. 149.—McDunnough, 1939, p. 35.
Prophtasia bistriatella (Hulst.) — Hampson, 1918, p. 106.
Saluria gemmatella (Hulst). — Hampson, 1918, p. 100.
Dracnosis.—This species is distinguished within the genus by the stalked veins M2;; coupled with the presence of two (subequal) cornuti in the vesica of the male.
DerscriptTion.—Frons conical, orange red; labial palpi with basal segments light reddish brown to white, second and third segments coral red on outer sides, light brown ventrally; maxillary palpi small, inconspicuous; antennae with scape coral red anteriorly and on inner side, white posteriorly and on outer side, shaft brown, red toward base, laminate in male; lateral regions of occiput, patagia, and tegulae orange red, vertex white behind antennae.
Forewings brownish red; Sc, R;, and R, white traced; R3, Ru, and M; black traced; A, white traced; M; and cubitals traced with varying mixtures of brown, pink, and white; A; orange traced; cell variable, often with more brown or orange than other areas; 11 veins; Re free from cell; M2,; stalked. Hindwings light brown, darker in apical area; 7 veins; M; and Cu, stalked.
Male genitalia with gnathos bearing short apical digitate projection, sometimes very short or absent. Juxta V-shaped. Valvae with costa bearing blunt tooth at apex, sparsely setose; sacculus minutely pubes- cent. Aedeagus with vesica bearing pair of subequal cornuti, smaller one serrate and more heavily sclerotized.
Female genitalia with ovipositor tip moderately setose, caudal margin finely setose. Posterior apophyses with base somewhat curved, spear-shaped; anterior very gradually thickened toward base.
REVISION OF PEORIINAE AND ANERASTIINAE 23
Typrs.—S. gemmatella and C. bistriatella, in the American Museum of Natural History; P. pamponeralla, in the U.S. National Museum.
Type patTa.—S. gemmatella, lectotype male, hereby designated, Illinois, collection G. D. Hulst, genitalia slide No. 3223, Carl Hein- rich, June 14, 1946; bestriatella, lectotype male, hereby designated, Colorado, collection G. D. Hulst, genitalia slide No. 3224, Carl Hein- rich, June 14, 1946; in the original description Hulst gives as locality and date: ‘“T'aken at San Bernardino, S. Cal., the last week in June’; pamponerella, lectotype male, hereby designated, Chimney Gulch, Golden, Colo., Aug. 12, 1907, Oslar; USNM 11854; genitalia slide No. 565, J. Shaffer, Nov. 23, 1965; lectoparatype: one male, same data as lectotype except no genitalia slide, F. Haimbach coll., Lot No. 72, Brackenredge Clemens Memorial collection, in the Academy of Natu- ral Sciences, Philadelphia.
SPECIMENS EXAMINED.—47 <’, 10 2 Distrisution (Map 6).—Colorado east to Iowa, Illinois, southern Ontario, New Jersey, northward along coast to Maine.
UNITED STATES: Connecticut: New Haven Co., East River, 19, July 13, 1909 (Charles R. Ely) [USNM];1 o, Aug. 12, 1909; 1 o, August 1911;1 #, July 10, 1912.
Intinors: Cook Co., Arlington Heights, 1c’, July 18, 1934 (A. L. McElhose) [CNHM]; 1 9, Aug. 15, 1936; Chicago, 1 9, Aug. 5, 1916 (E. Beer) [CNHM]; Macon Co., Decatur, 1 &, Aug. 1, 1890 (W. Barnes) [USNM]; Hancock Co. Webster, 1 o&, Aug. 12, 1883 [USN M].
InpIANA: Lake Co., Hessville, 1 9, July 15, 1905 (A. Kwiat) [CNHM];1 @, Aug. 1, 1908 (E. Beer) [CNHM].
Towa: Story Co., Ames, 1 o&, June 27, 1912 (G. C. Decker) [USNM]; 1 @, July 17, 1932; 1 9, August [USNM]; Woodbury Co., Sioux City, 1 o, June 25, 1936 (C. N. Ainslie) [UM].
Marne: York Co., Kennebunk, 1 o, Aug. 11, 1901 (G. H. Clapp) [CM]; 1 of, Aug. 22, 1901.
MassacuuseEtts: Nantucket Co., Nantucket, 1 o&, July 21, 1945 (C. P. Kimball) [CPK]; Barnstable Co., Barnstable, 1 o&, Aug. 19, 1952 (C. P. Kimball) [CPK]; Woods Hole, 3 @, August 1917 (W. T. M. Forbes) [CU]; Plymouth Co., Agric. Exp. Sta., East Wareham, 1 o’, Aug. 7, 1964 [CPK]; 1 o, Aug. 22, 1964.
New Jersey: Burlington, Brown’s Mills, 1 @, August 31, (F. H. Benjamin) [CU]; New Libson, 1 o, June 12, 1942 (E. P. Darlington) [ANS]; 1 o, Aug. 4, 1942; Whitesbog, 1 o&, June 22, 1940 (E. P. Darlington) [ANS]; 1 o&, Aug. 5 1940; 1 o&, Sept. 19, 1936; Ocean Co., Lakehurst, 1 9, Aug. 30, 192? (F. M. Schott) [USNM]; Lakehurst, Wrangle Brook Rd., 2 o', June 27, 1954 (J. G. Franclemont) [JGF]; 2 o&, June 27, 1955; 1 o&, Aug. 9, 1955.
New York: Suffolk Co., Orient, 1 @, July 16, 1932 (Roy Latham) [CU]; 1 o, Apr. 21, 1935 [AMNH]; 1 9, June 14, 193?, [CU]; 1 9, July 3, 1945; 1 9, Sept. 4, 1945; 1 o, Sept. 16, 1945; 1 o&, July 7, 1947 [AMNH]; 2 &, Aug. 17, 1947; 1 o, no date [CNC].
CANADA: Ontario: Kent Co., Chatham Lab., 1 o&, Aug. 28, 1932 [CNC]; 1 o, July 12, 1934; 1 & , July 18, 1934; 1 &, July 20 ,1935.
24 U.S. NATIONAL MUSEUM BULLETIN 280
LOcALITY UNKNOWN: Edgebrook (Prob. Cook Co., Illinois), 1 @, July 8, 1911 (A. Kwiat) [USN M]. 2 3, no date, [INHS]; 2 o, 1 9, no date [USNM].
Peoria roseotinctella (Ragonot), new combination Ficures 10, 61, 77, 112, 150
Statina roseotinctella Ragonot, 1887, p. 19; 1889, p. 117. — Hulst, 1890, p. 216 — Smith, 1891, p. 85. — Ragonot, 1901, p. 416. — Hulst, 1902, p. 440. — Barnes and McDunnough, 1917, p. 150. — Grossbeck, 1917, p. 134. — Hampson, 1918, p. 60. — McDunnough, 1939, p. 36. — Kimball, 1965,
. 251.
Calera punctilimbella Ragonot, 1888, p. 50. — Hulst, 1890, p. 217. — Smith, 1891, p. 85. — Ragonot, 1901, pp. 417-418. — Hulst, 1902, p. 441. — Barnes and McDunnough, 1917, p. 150. — Hampson, 1918, p. 59. — Forbes, 1923, p. 639. — McDunnough, 1939, p. 36. [New synonymy.]
Statina bifasciella Hampson im Ragonot, 1901, pp. 416-417. — Barnes and McDunnough, 1917, p. 150. — Hampson, 1918, p. 60. — McDunnough 1939, p. 36. [New synonymy.]
Dracnosts.—The transverse posterior line of dots on the forewing veins delimits this species within the genus.
DescripTion.—Frons conical, deep red, often brown laterally; labial palpi with basal segments white, second and third segments deep red on outer sides, ventral third lighter, dorsal third often brown; maxillary palpi moderately small; antennae with scape deep red on inner side, white on outer side, shaft light brown, pink near base, laminate in male; occiput, patagia, tegulae, vertex, and dor- sum of thorax deep red; legs pink on outer sides, light brown on inner sides, tarsi often brown.
Forewings orange red, mixed with white anterior to cell; Sc and Rs sometimes white traced, Rs often bordered posteriorly by a broad line of scattered black scales extending from base to apex; transverse posterior indicated by black spots on Rs, My, M3, Cui, Cue, and Ag; terminal line indicated by dark spots on M3, Cui, Cus, and A, fold; terminal spots visible on lower surface of wing; 10 veins; R» stalked with Ris, Me fused with M;; M; stalked with Cu;. Hindwings with 6 veins; M; and Cu, completely fused.
Male genitalia with medial process of uncus bearing small apical bulge. Gnathos weakly sclerotized, without apical or lateral processes, occasionally with very short apical process. Juxta U-shaped. Valvae with costa sparsely setose, terminating in a dorsally directed hook; sacculus densely pubescent. Aedeagus with vesica bearing equal pair of finely serrate cornuti.
Female genitalia with ovipositor moderately setose laterally, caudal margin with numerous fine setae. Posterior apophyses straight, base flat and spearlike; anterior curved, tapering gradually to slightly thickened base.
REVISION OF PEORIINAE AND ANERASTIINAE 25
Typrs.—S. roseotinctella, in the Museum National d’Histoire Naturelle; C. punectilimbella, in the Zoologisches Museum der Hum- bolt-Universitit, Berlin; S. bcfasciella, in the British Museum (Natural History).
Type pata.—s. roseotinctella, lectotype male, hereby designated, Florida, genitalia slide No. 539, J. Shaffer, June 20, 1965.
C. punctilimbella, lectotype female, hereby designated, Carolina; genitalia slide No. 818, J. Shaffer, Mar. 20, 1967.
S. bifasciella, lectotype female, hereby designated, labeled as follows: “11/9; Type; Belfrage Texas 1869; Stainton Coll. 93—134.; Statina bifasciella type 2 Hmpsn; @ genitalia slide I-4-1967 J. Shaffer No. “02-”
SPECIMENS EXAMINED.—57 co’, 259.
DistrisuTIon (Map 6).—Kansas and eastern Texas to Florida, southern Florida north to New Jersey.
UNITED STATES: AtasBama: Macon Co., LaPlace near Tuskeege, 4 oc, June 9, 1917 [CU].
District or CoLtumBIA: Washington, 19, June 1902 (Aug. Busck) [USN M].
Fioripa: Alachua Co., Gainesville, 19, June 2, 1927 (J. Speed Rogers) [CU]; 1 o, June 3, 1927; 19, June 29, 1927; 1 o&, July 7, 1927; 13 o&, July 8, 1927; 3 o, July 10, 1927; 3¢, July 1927; 19, Apr. 24, 1952 (O. Peck) [CNC]; 16, Sept. 3, 1956 (H. A. Denmark) [CPK]; no locality, 167, Sept. 13, 1956 (H. A. Denmark) [CPK]; Dade Co., Homestead, 19, Aug. 7, 1963 (D. O. Wolfenbarger) [CPK]; 1¢, Mar. 10, 1964; 1c’, 19, Sept. 29, 1964; Princeton, 19, Apr. 4, 1952 (J. R. Vockeroth) [CNC]; Escambia Co., Pensacola, 1c’, May 22, 1961 (Shirley Holls) [CPK]; Hernando Co., Weeki Wachee Springs, 1c’, May 28, 1960 (J. F. May) [CPK]; Highlands Co., Archbold Biol. Sta., 16, June 27, 1964 (Jay C. Shaffer) [JCS]; Hillsborough Co., Stemper, 1c, Aug. 19, 1912 (G. Krautwurm) [CM]; 1<7, Sept. 1 1912; 16, Sept. 6, 1912; 167, Oct. 3, 1912; 16, Oct. 6, 1912; 2¢, Oct. 30, 1912; Manatee Co., Oneco, 16, Aug. 3, 1953 (Paula Dillman) [CPK]; 1, Mar. 28, 1957 (J. G. Franclemont) [JGF]; Orange Co., Orlando, 14, June 29, 1927 (C. C. McBride) [CU]; 14, July 24, 1927; Winter Park, 19, July 4, 1942 (H. T. Fernald) [USN M]; Volusia Co., Cassadaga, 1c’, Oct. 18, 1963 (S. V. Fuller) [CPK]; Wakulla Co., Panacea, 2¢°, Aug. 11, 1926 (C. O. Handley) [USN M].
Groraia: Bryan Co., Clyde, 1, Sept. 11-12, 1931 (Bradley and Knorr) [C U}.
Kansas: Riley Co., Manhattan, 19, June 11, 1933 (H. L. Nonamaker) [KS U].
LovuisiaANa: Vernon Ph., 19, August (G. Coverdale) (USN M].
Misstssipri1: Forrest Co., Camp Shelby, 306’, 29, Oct. 1-15, 1944 (C.D. Michener) [AMNH]; Hinds Co., Clinton, 19, June 12, 1960 (Bryant Mather) [BM].
New Jersry: Ocean Co., Lakehurst, Wrangle Brook Rd., 1c, June 30, 1965 (Jay C. Shaffer) [JCS].
Norrs Carona: Polk Co., Tryon, 14, Aug. 8, 1904 (Fiske) [USNM]; 2¢, Aug. 9, 1904; 16, Aug. 10, 1904; 16, Aug. 11, 1904; 167, no date.
Sour Caroxina: Oconee Co., Cherry Hill Rec. Area, Route 107 (2000 ft.), 13, Aug. 22, 1958 (J. G. Franclemont) [JGF].
26 U.S. NATIONAL MUSEUM BULLETIN 280
TENNESSEE: Knox Co., Knoxville, 19, June 26, 1916 (G. G. Ainslie), [USN M]. Texas: Blanco Co., 19, no date [USNM]; Liberty Co., Devers, 17, 29, June 21, 1917 [CU]; Nueces Co., Corpus Christi, 69, Sept. 25-Oct. 15, 1943 (W. M. Gordon) [CU].
Peoria johnstoni, new species Fieurss 1, 78, 113, 151
DraGnosis.—Among the three other species of Peoria with similar venation, johnstoni differs from approximella and luteicostella in lacking a well-developed white band anterior to the cell, and from bipartitella in having the forewings a rather uniform brownish orange. Either the presence of two cornuti on the vesica of the aedeagus or the absence of medial or lateral processes on the gnathos will de- limit males of johnstoni from the above three species.
Description.—Frons conical, reddish brown; labial palpi with basal segments white, second and third segments reddish brown dorsally and on outer sides, lighter ventrally and on inner sides; maxillary palpi small; antennae light brown, scape light pink an- teriorly, male subserrate; occiput, patagia, and tegulae reddish brown, vertex somewhat lighter.
Forewings brownish orange; Sc, radials, cubitals, M3, and Az white traced, costa traced with narrow inconspicuous white line; brownish orange on lower surface. Venation as in P. bipartitella.
Male genitalia with medial process of uncus bearing small ventral bulge at apex. Gnathos weakly sclerotized, without apical or lateral processes. Juxta U-shaped. Valvae with costa sparsely setose; sacculus densely setose. Aedeagus with vesica bearing pair of ovate subserrate cornuti.
Female genitalia with ovipositor moderately setose along caudal margin. Posterior apophyses straight, base flat and shaped like parallelogram with short side horizontal; anterior slightly curved, tapering gradually to thickened base.
Typr.—In the Canadian National collection.
Typre pata.—Holotype, male, Fort Davis, Jeff Davis County, Tex., May 20, 1950, E. C. Johnston; C.N.C. Type No. 9440; genitalia slide No. 591, J. Shaffer, Mar. 24, 1966.
Paratypes: Five males, same data as holotype, no genitalia slides prepared.
SPECIMENS EXAMINED.—160', 159.
DistriputTion (Map 7).—Mexican Highland Province; southern Arizona and New Mexico to southwestern Texas.
UNITED STATES: Arizona: Pima Co., Baboquivari Mts., 39, Sept. 1-15,
1923 (O. C. Poling) [USNM]; 19, Oct. 1-15, 1923; 19, Apr. 15-30, 1924; 3¢7 29, Oct. 1-15, 1924; 16, 29, Oct. 15-30, 1924; Pinal Co., Oracle, 1¢, June 5,
REVISION OF PEORIINAE AND ANERASTIINAE 27,
1935 (Grace H. and John L. Sperry) [AMNH]; Santa Cruz Co., Pena Blanca (3950 ft.), 1c, May 31, 1963 (J. G. Franclemont) [JGF]; county unknown (southeastern Arizona), Huachuca Mis., 1o’, June 1, 1935 (J. A. Comstock) [CNC]; 19, May 28, 1935 (Grace H., and John L. Sperry) [CNC]; 19, June 1, 1935 [AMNH].
New Mexico: Eddy Co., White City, 1, May 15, 1950 (E. C. Johnston) [CNC].
Texas: Brewster Co., Alpine, 49, May 22, 1950 (E. C. Johnston) [CNC]; Jeff Davis Co., Limpia Canyon, 8c, May 20, 1950 (E. C. Johnston) [CNC].
Peoria santaritella (Dyar), new combination
Fiaures 5, 79, 114, 152
Ollia santaritella Dyar, 1904, p. 108.—Barnes and McDunnough, 1917, p. 149.— MecDunnough, 1939, p. 36.
Diacnosis.—The forewings are white anterior to the cubitus, red between the cubitus and the A, fold, and light pink posterior to the fold; the combination of these features serves to delimit the species within the genus.
Description.—Frons conical, blood red; labial palpi with basal segments white, second and third segments red on outer sides, white ventrally and on inner sides; maxillary palpi small; antennae with scape red, shaft brown, male serrate and fasciculate; occiput deep red behind eye, vertex posterior to antennae and dorsal area of occiput white, patagia and tegulae pink; legs white, tarsi light brown, forelegs pink on inner sides.
Forewings white anterior to cubitus, a few scattered red scales anterior to cell, costal margin bordered with red near base; red between cubitus and A, fold, bounded distally by M3; and Cuz; light pink posterior to red line; underside with red showing through faint- ly; 10 or 11 veins; Re» free from cell; M2;, stalked for about half its length or completely fused, both conditions about equally common, sometimes (rarely) differing on right and left wings of same specimen. Hindwings with M; and Cu, stalked.
Male genitalia with gnathos bearing short apical digitate projection. Juxta V-shaped. Valvae with costa terminating in a blunt tooth, cucculus not projecting beyond costa; sacculus with fine hairs. Aedeag- us with vesica bearing single serrate cornutus.
Female genitalia with ovipositor tip sparsely setose, caudal margin with numerous fine setae. Posterior apophyses with base tapering to point, not flattened or broadened, curved near base; anterior tapering gradually to slightly thickened base.
Typr.—In the U.S. National Museum.
Typr pata.—Holotype male, Santa Rita Mountains, Arizona, “7,6,” E. A. Schwarz; USNM 7893; genitalia slide No. 569, J. Shaffer,
285-934— 683
28 U.S. NATIONAL MUSEUM BULLETIN 280
Nov. 23, 1965. In the original description Dyar gives the date of capture as June 7.
SPECIMENS EXAMINED.—4 0’, 21 9.
DistrisuTIon (Map 7).—Northern extension of Sierra Madre Occidental into southeastern Arizona.
UNITED STATES: Arizona: Cochise Co., Paradise, Chiricahua Mts., 19, July 3, 1954 (Cazier and Gertsch) [AMNH]; Southwestern Res. Sta. (5400 ft.), 5 mi. west Portal, 19, May 28, 1956 (Cazier and Ordway) [AMNH]; 19, May 25, 1958; 49, June 11, 1958 (W. J., and J. W. Gertsch) [AMNH]; 29, June 12, 1958; 19, June 16, 1958 (M. A. Cazier) [AMNH]; 19, May 27, 1960 (Carl W. Kirkwood) [CPK]; 19, June 2, 1960; 1, May 10, 1961 (Gertsch and Cazier) [AMNH]; Pima Co., Santa Rita Mts., Madera Canyon (4400 ft.), 19, June 12, 1963 (J. G. Franclemont) [JGF]; Santa Cruz Co., Santa Rita Mts., Madera Canyon (4800 ft.), 19, June 19, 1960 (David A. Wallesz) [CU]; (5800 ft.), 19, June 24, 1960; Madera Canyon (4800 ft.), 1, May 26, 1963 (J. G. Franclemont) [JGF]; 1%, 19, June 19, 1963; 19, June 13, 1960; 19, June 3, 1963; (5600 ft.), 16, June 11, 1963; 12, June 18, 1963; Pena Blanca (3950 ft.), 12, June 7, 1963 (J. G. Franclemont) [J GF]; county unknown, Huachuca Mts., 19, June 2, 1935 (Grace H., and John L. Sperry) [AMNH].
Peoria holoponerella (Dyar), new combination Fiaurss 3, 4, 80, 115, 153
Ollia holoponerella Dyar, 1908, p. 117. — Barnes and McDunnough, 1917, p. 149. — McDunnough, 1939, p. 36.
Diaanosis.—The light buff forewing with its conspicuous white band anterior to the cell delimits this species within Peoria. A similar white band marks the forewings of approximella and luteicostella, but in both of these species the forewings are marked with red and lack vein Mp.
DescripTion.—Frons conical, brown; labial palpi with basal segments white, second and third segments brown on outer sides, light brown to white ventrally and on inner sides; maxillary palpi small; antennae silver, male shaft serrate and fasciculate; occiput behind eye, patagia, and tegulae light brown, vertex and dorsum of occiput white; legs white, tarsi brown.
Forewings with white band anterior to cell, bordered anteriorly near base with dark brown, posteriorly by black line; ground light buff; veins traced with black scales. Hindwings light brown. Venation as in gemmatella.
Male genitalia with gnathos bearing medial digitate projection. Juxta V-shaped. Valvae with costa unarmed, bearing scattered setae; sacculus with fine hairs, Aedeagus with vesica bearing single round serrate cornutus.
Female genitalia with ovipositor slender, caudal margin rather heavily setose, bare laterally, Apophyses straight, posterior hooked upward and coming to point at base; anterior with thickened base.
REVISION OF PEORIINAE AND ANERASTIINAE 29
Typr.—In the U.S. National Museum.
Type pata.—Holotype male, San Bernardino Ranch, Cochise County, Ariz., 3750 ft., August, F. H. Snow; USNM 11855; genitalia slide No. 568, J. Shaffer, Nov. 23, 1965.
SPECIMENS EXAMINED.—18<7, 29.
DistriBuTion (Map 7).—Southwestern United States.
UNITED STATES: Arizona: Cochise Co., Douglas, 2 &, August (F. H. Snow) [UK]; San Bernardino Ranch (3750 ft.), 5 #@, 1 9, August (F. H. Snow) [UK].
Cauirornia: Inyo Co., Furnace Creek, Death Valley, 3 o&, Apr. 21, 1942 (George Willett) [LACM]; Triangle Springs, Death Valley, 1 9, Apr. 14-15, 1942 (George Willett) [LACM]; 1 o, Apr. 17, 1942.
New Mexico: Sandoval Co., Jamez Springs (6200 ft.), 4 o&, July 16, 1950 (T. Cohn, P. Boone, M. Cazier) [AMNH].
Texas: Jeff Davis Co., Limpia Canyon, 2 @, May 20, 1950 (E. C. Johnston) [CNC].
Uran: Locality unknown, 1 o’, July 1900 (Poling) [USNM].
Discussion.—The five specimens from Death Valley entirely lack
brown or black markings on the forewings.
Peoria approximella (Walker) Figures 14, 59, 81, 116, 154
Eurhodope approzimella Walker, 1866, p. 1722.
Anerastia haematica Zeller, 1872, pp. 555-556. — Grote, 1879, p. 12, pl. 2, fig. 14
Nephopteryz roseatella Packard, 1873, p. 270.
Peoria haematica (Zeller). — Ragonot, 1889, p. 117. — Hulst, 1890, pp. 213-214. — Smith, 1891, p. 85.
Peoria approximella (Walker). — Ragonot, 1901, p. 386. — Hulst, 1902, p. 439. — Barnes and McDunnough, 1917, p. 149. — Grossbeck, 1917, p. 134. — Forbes, 1923, p. 638. — McDunnough, 1939, p. 36. — Kimball, 1965, p. 251.
Hypsotropa approximella (Walker). — Hampson, 1918, p. 77.
Hypsotropa cremoricosta Hampson, 1918, p. 74. — McDunnough, 1939, p. 36. [New synonymy.]
Draenosis.—The presence of both a prominent white band anterior to the cell and a fainter one posterior to vein Az marks this species as distinct within the genus. The gnathos is also unique in that it bears a pair of short lateral finger-like processes.
Description.—Frons conical, blood red to orange red; labial palpi with basal segments varying white to light pink, second and third segments somewhat rough scaled, blood red to orange red dorsally and on outer sides, usually paler ventrally and on inner sides; maxillary palpi small; antenna with scape red on anterior and inner sides, light pink to white on posterior and outer sides, shaft light brown, pink toward base, laminate in male; occiput laterally, patagia, and tegulae orange red, vertex light brown to white.
Forewings with broad white band anterior to cell, narrowing to point just short of apex, bordered anteriorly with black near wing
30 U.S. NATIONAL MUSEUM BULLETIN 280
base, bordered posteriorly with black, fading into blood red triangular area extending to anal vein; brownish white posterior to anal vein; anal and costal bands joined near wing base; fringe white. Venation as in bipartitella.
Male genitalia with gnathos weakly sclerotized, bearing pair of short, lateral, subapical projections. Juxta scoop-shaped. Valvae with costa terminating in free spine not attaining apex of bluntly rounded cuc- culus; sacculus minutely pubescent. Aedeagus with vesica unarmed.
Female genitalia with ovipositor tip moderately setose, caudal mar- gin with numerous fine setae, sparsely setose laterally. Apophyses uni- form, curved at base; posterior tapering to point at base, base not flattened.
Typres.—E. approximella, A. haematica, and H. cremoricosta in the British Museum (Natural History); N. roseatella, in the Museum of Comparative Zoology (Harvard University).
Type pata.—EL. approzimella, Walker reports: ‘‘North America. From Mr. Carter’s collection.”’ Type examined by Mr. Paul Whalley.
A. haematica, Zeller in his description noted: “. .. New York und aus Massachusetts ....’’ Type examined by Mr. Paul Whalley.
N. roseatella, two specimens, both from Massachusetts, and both Type No. 14282, one from Dorchester, the other hereby designated as lectotype and labeled male genitalia slide No. 540, J. Shaffer, June 24, 1965. In his description Packard indicated that he had two males and noted: ‘Length of fore wing .37 of an inch. Dorchester, Mass. (F. G. Sanborn).”
H. cremoricosta, lectotype male, hereby designated, labeled as fol- lows: ‘Type; Colorado Springs, Colorado. 97-278 June-Aug 1896; Hypsotropa cremoricosta type o'. Hmpsn.; o& genitalia slide I-4—1967 J. Shaffer No. 703.”
SPECIMENS EXAMINED.—562 <7, 302 9.
DistriputTion (Map 5).—Southern Canada west to Alberta, south- ward in mountains to Utah and New Mexico, east to Tennessee and Georgia, north to Nova Scotia; absent from Florida and the Gulf Coastal Plain.
UNITED STATES: Cotorapo: El Paso Co., Rock Creek Canyon, Colorado Springs, 2 o#, Aug. 10, 1957 (Margot May) [CPK];1 o&, Aug. 18, 1957; 4 @#, Aug. 14, 1957; 1 o&, Aug. 23, 1957; 2 o, June 27, 1960; 1 &, June 28, 1960; 1 o, July 15, 1960; 1 o, July 9, 1961; 1 o&, July 19, 1961; 6 &, June 30, 1963; 2 7, July], 1963; 1 o, July 3, 1963; 2 o, July 14, 1963; 1 o, July 17, 1963; 1 o, July 18, 1963; 1 o&, Aug. 11, 1963; 1 o&, June 27, 1964; 3 @, June 28, 1964; vic. Colorado Springs, 1 ?, July 7, 1939 [AMNH]; Jefferson Co., Chimney Gulch, Golden,1 &, Sept. 3, 1908 (Oslar) [ANS]; Larimer Co., Estes Park, 1 o&, Aug. 29, 1936 (Grace H. and John L. Sperry) [AMNH]; Teller Co., Big Springs Ranch, Florissant, 1 <, July 13, 1960 (Thomas C. Emmel) [LACM]; 1 0, July 21, 1960; 2 &, July 23, 1960; 2 o, July 26, 1960; 1 o&, July 29, 1960; 1 o&, July 30, 1960; 1 o, July 31, 1960; 2 o&, Aug. 2, 1960; 1 o, Aug. 3, 1960, 1 &, Aug. 4, 1960; 1 o, Aug. 8, 1960;
REVISION OF PEORIINAE AND ANERASTIINAE 31
1 o, Aug. 11, 1960; 1 o&, Aug. 12, 1960; 1 o, Aug. 14, 1960; locality unknown, 1 o', no date [AMNH].
Connecticut: Windham Co., South Shore, Killingly Pd., 1 9, July 25, 1930 (A. B. Klots) [AKB]; Putnam, 1 o, Aug. 14, 1954 (A. B. Klots) [ABK]; 1 ¢, July 1, 1960; 1 9, July 20-25, 1961; 1 9, July 25-30, 1961.
District or CoLumsBia: 1 o’, Aug. 17, 1899 (August Busck) [USNM]; 2 9, July 1901; 2 9, June 1902; 1 9, June 1907; 1 9, June.
Georges: Charlton Co., 1 9, June 14, 1946 (Otto Buchholz) [ANS].
Inuino1s: Cook Co., Arlington Heights, 1 @, June 20, 1930 (A. L. McElhose) [CNHM]; 1 9, June 23, 1930; 1 9, June 24, 1930; 1 9, July 29, 1930; Chicago, 3 o’, June 1900 [USNM];1 9, July 14, 1904; 1 &, July 8, 1906 (W. J. Gerhard) [CNHM]; 1 ao, June 1912 (A. Kwiat) [CNHM]; Jackson Co., Murphysboro, 1 9, Aug. 10, 19382 (W. J. Gerhard) [CNHM]; McHenry Co., Algonquin, 1 ¢&, July 6, 1903 (W. A. Nason) [INHS]; 1 9, July 16, 1904; 1 o&, June 8, 1906; McLean Co., Normal, 1 9, June 19, 1884 [INHS]; Macon Co., Decatur, 1 ¢& June 8-15 (Barnes) [USNM]; Putnam Co., 1 o&, May 31, 1934 (M. O. Glenn) [MOG]; 1 o, June 21, 1954; 1 o, Sept. 5, 1955; 1 o, June 8, 1959; 1 9, July 19, 1962; 1 9, June 25, 1964; 1 9. July 15, 1964; Magnolia, 1 6, June 29, 1932 (Murray O. Glenn) [USNM]; locality unknown, 2 o’, no date [USNM].
InpDIANA: Lake Co., Hessville, 1 o, May 14, 1905 (A. Kwiat) [CNHM]; 1 o',; June 17, 1905; 1 o, June 17, 1912.
Iowa: Johnson Co., Iowa City, 1 9, July 12, 1898 (Wickham) [AMNH]; 1 9, July 27, 1898; Story Co., Ames, 1 o, July 9, 1915 (G. G. Ainslie) [USNM]; Woodbury Co., Sioux City, 1 9, July 23, 1923 (C. N. Ainslie) [UM]; locality unknown, 1 <a’, no date (C. P. Gillette) [USN M].
Kansas: Pottawatomie Co., Onaga, 1 o, June 17, 1901 (Crevecoeur) [KS]; 1 6, July 7, 1901; Riley Co., 1 o&, June 22 (F. Marlatt) [KS]; 1 9, July; 1 0, July 28.
Kentucky: Rockeastle Co., 1 o&, June 24, 1955 (R. Beebe) [CNC].
Marne: Franklin Co., Rangeley, 1 o, June 23, 1938 (V. H. dos Passos) [AMNH]; 1 o&, July 9, 19388; Knox Co., Rockport, 1 &, July 10, 1941 (Morgan Hebard) [ANS]; Penobscot, Orono, 2 ?, no date, [CU]; York Co., Kennebunk- port, 2 o, 1 2, July (G. H. Clapp) [CM]; 3 0, 1 9, August.
Maryuanp: Montgomery Co., Plummer Isl., 1 9, May 1903 (August Busck) [USNM]; 1 &, June 1903; 1 9, Aug. 2, 1905 (H. S. Barber) [USNM]; 2 9, Aug. 8, 1905 (Barber and Schwarz) [USNM]; Prince Georges Co., Beltsville, 1 o’, June 10, 1939 (L. J. Bottimer) [ANS].
MassacuvuseEtts: Barnstable Co., Woods Hole, 1 9, August 1917 (W. T. M. Forbes) [CU]; Barnstable, 1 o, July 5, 1949 (C. P. Kimball) [CPK]; 1 o, July 6, 1949; 4 o&, July 7, 1949; 1, June 24, 1958; 1 o&, Aug. 1, 1958; 1 9, June 18, 1962; West Barnstable, 3 o, July 15, 1949 (C. P. Kimball) [CPK]; 1 &, July 16, 1949; Dukes Co., Martha’s Vineyard, 1 #, June 17, 1931 (F. M. Jones) [ANS]; 1 9, July 22 [(CPK]; Essex Co., Magnolia, 1 @, June 29, 1900 (Holland) [CM]; Hamp- den Co., Chicopee, 1 9, no date (F. Knab) [USNM]; Nantucket Co., Nantucket, 1 o&, July 9, 1941 (C. P. Kimball) [CNC]; Plymouth Co., Agric. Exp. Sta., East Wareham, 2 o’, Aug. 7, 1961 [CPK]; 1 o&, July 3, 1962; 1 &, July 19, 1962; 1 o, Aug. 17, 1964; 1 o&, Aug. 18, 1964; 1 o&, Aug. 20, 1964; Suffolk Co., Arling- ton, 1 o, June 25, 1920 (C. S. Anderson) [BPI]; Boston, 1 °, no date [AMNH]; Worcester Co., Paxton, 2 &, July 3, 1945 (W. T. M. Forbes) [CU]; locality unknown, 1 o’, June 27, 1868 (BM, ex Zeller coll.].
Micuiean: Kalamazoo Co., Gull Lake Biol. Sta., 1 9, July 21, 1959 (Roland L. Fischer) [CNC].
5) U.S. NATIONAL MUSEUM BULLETIN 280
Minnesota: Cass Co., 1c’, June 29, 1937 (L. W. Orr), [UM]; 1c’, June 30, 1937; 10, July 2, 1937; 16’, July 11, 1937; Cass Lake, 1’, June 18, 1934 (A. A. Granovsky) [UM]; 19, July 17, 1936 (R. H. Daggy) [UM]; 1c’, Aug. 3, 1936; 1, Aug. 14, 1986; Hennepin Co., Saint Anthony Park, 19, July 12, 1900 [UM]; Olmsted Co., 19, no date [UM]; Polk Co., Crookston, 14, July 18, 1935 (D. G. Denning) [UM]; 10’, June 27, 1937; 167, July 4, 1937; 1¢, July 23, 1939 (A. W. Buzicky) [UM]; Crookston, Red Lake River, 1, July 9, 1935 (D. G. Denning) [UM]; Pope Co., Sedan, 1<, July 7, 1927 (D. Denning) [UM]; Ramsey Co., St. Paul, 14, July 21, 1927 (Carl T. Schmidt) [UM]; county unknown, Itasca Park, LaSalle Valley, 1c’, June 21, 1940 (C. E. Mickel) [UM]; 1c’, June 23, 1940; 1, June 24, 1940; 10, July 7, 1940; 2¢°, 19, July 9, 1940; 10, July 12, 1940; 1%, July 14, 1940.
Missouri: Clinton Co., Lathrop, 19, June 22, 1955 (E. C. Becker) [CNC]; city of St. Louis, 567, 59, June 1904 (August Busck) [USNM]; 1¢, July 1904; 14,19, Aug. 6, 1906 (McElhose) [CNHM].
NEBRASKA: Sioux Co., Canon region north of Harrison, 40, July 19, 1917 (R. A. Leussler) [USNM, AMNH].
New Hampsuire: Coos Co., Randolph, 19, Aug. 16, 1936 (Grace H. and John L. Sperry) [AMNH]: Grafton Co., Franconia, 2¢°, no date [AMNH]; Rockingham Co., Hampton, 1o’, June 27, 1913 (S. Albert Shaw) [USNM]; 19, July 18, 1914.
New Jersey: Bergen Co., Bergenfield, 19, July 3 (F. M. Schott) [AMNH]; Oakland, 19, July 25, 1947 (C. P. Kimball) [CPK, CNC]; 19, July 26, 1947; 19, July 27, 1947; 16%, Aug. 8, 1948; 16, Aug. 9, 1948; Aug. 10, 1948; 1¢, Aug. 12, 1948; 167, Aug. 18, 1948; Ramsey, 16°, 39, July 12, 1935 [AMNH]; 1%, July 29, 1935 [AMNH]; Burlington Co., New Lisbon, 1¢, July 14, 1930 (E. P. Darlington) [ANS]; 16, July 1, 1932; 19, July 19, 1932; 1¢, July 29, 1931; 1o, July 20, 1942; Essex Co., Montclair, 1¢, June 15 (W. D. Kearfott) [USNM]; 1¢, July 10; 16, July 18; Newark, 1¢, June 13, 1898 [USNM]; Middlesex Co., Dayton, 20’, June [USNM]; New Brunswick, 1¢, June 11 [AMNH]; 1¢, June 1933 [AMNH]; Monmouth Co., Como, 19, July 1919 [AMNH]; Morris Co., Mendham, 1.7, 19, June 20, 1934 [AMNH]; Ocean Co., Wrangle Brook Road, Lakehurst, 1o¢, June 17, 1955 (J. G. Franclemont) [J GF]; Union Co., Elizabeth, 19, ‘‘8-13’”” [CNHM]; county unknown, 19, no date [AMNH].
New Mexico: McKinley Co., McGaffey, Zuni Mts. (7500 ft.), 1¢, July 22, 1962 (E. and I. Munroe) [CNC]; 2¢, July 23, 1962; Sandoyal Co., Frijoles Canyon, Bandelier Nat. Mon. (6050 ft.), 1, July 18, 1962 (E. and I. Munroe) [CNC]; county unknown, 1’, June 30, 1935 (Grace H. and John L. Sperry) [AMNH].
New York: Cattaraugus Co., Otto, 1 9, July 17, 1882 [CU]; Clinton Co., Peru, 20°, 19, June 21, 1915 [CU]; 19, July 1, 1916; 10, July 2, 1916 (Mix and Everett) [CU]; Delaware Co., 4c, Aug. 3, 1962 (Robert Silberglied) [JCS]; 43, 9, Aug. 5, 1962; Erie Co., East Aurora, 14, July 20, 1912 (E. Wild) [CU]; North Collins, 1¢@, 29, June 27, 1939 (W. T. M. Forbes) [CU]; 17, June 28-29, 1939; Sardinia, 1o, July 24, 1940 (L. R. Rupert) [CU]; 19, July 27, 1940; Jefferson Co., Picton Island, Clayton, 1, June 22, 1958 (B. Heineman) [AMNH]; Lewis Co., 1, June (W. W. Hill) [UK]; Monroe Co., 19, July 11, 1945 [CNC]; 1, June 27, 1947 [CNC]; 19, July 9, 1947 [CNC]; 19, July 11, 1947 [CNC]; 10, July 21, 1947 [CPK]; 19, June 28, 1948 [CNC]; 19, June 29, 1948 [CNC]; 10, July 1, 1948 [CPK]; Rochester, 14, June 26, 1932 [ABK]; New York City, West Farms, 19, no date (J. Angus) [AMNH]; Orange Co., Tuxedo, 19 , June 28, 1928 [AMNH]; 19, July 10, 1928 [AMNH]; Oswego Co., Minetto, 14, June 19,
REVISION OF PEORIINAE AND ANERASTIINAE 33
1938 (W. T. M. Forbes) [CU]; 2c7, 29, June 21, 1938; 36, 99, June 22, 1938; 2c, 89, June 24, 1938; 19, Aug. 22, 1938; Otsego Co., Oneonta, 19, July 4, 1935 (H. K. Townes) [CU]; 19, July 5, 1935; Saint Lawrence Co., Oswegatchie, 3c, Aug. 3, 1926 (B. K. Smith) [CU]; Suffolk Co., Flanders, 16, Sept. 1, 1946 (Roy Latham) [CU]; Huntington, 19, July 25, 1925 (F. M. Schott) [USNM]; Montauk, 19, Sept. 8, 1946 (Roy Latham) [CU]; Orient, 19, July 28, 1931 (Roy Latham) [CU]; 1c’, July 6, 1932; 16, Aug. 3, 1932; 19, Aug. 4, 1932; 16, Nov. 6, 1932; 16, June 22, 1934; 14%, June 22, 1946 (Roy Latham) [AMNH]; 16’, June 21, 1947; 19, June 23, 1947; 1¢, July 21, 1947; 1¢, June 12, 194?; Shelter Island, 19, Sept. 14, 1946 (Roy Latham) [CU]; Southold, 19, July 1, 1934 (Roy Latham) [CU]; 1, July 24, 1946; Tompkins Co., Ithaca, 2.7, July 4, 1882 [CU]; 30°, 69, July 8, 1882 [CU]; 14, July 14, 1885 (E. H. Sargent) [CU]; 1°, June 13, 1893 [CU]; 19, July 7, 1893 [CU]; 14, June 1913 [CU]; 19, July 2, 1914 [CU]; 1°, June 26, 1915 [CU]; 14, June 15, 1919 [CU]; 1.7, June 16, 1919 [CU]; 19, June 21, 1919 [CU]; 29, June 21, 1922 [CU]; 1c, June 29, 1922 [CU]; 29, June 30, 1922 [CU]; 1c’, 19, July 2, 1922 [CU]; 19, July 4, 1944 [CU]; 16’, July 7, 1922 [CU]; 16, July 2, 1925 [CU]; 10, July 2, 1925 [CU]; 19, July 9, 1925 [CU]; 19, July 25, 1926 (Bolton K. Smith) [CU]; 19, July 5, 1928 [AMNH]; 1c, 19, June 16, 1929 [CU]; 14, 19, July 15 [CU]; 29, June 16, 1939 [CU]; Six Mile Creek, Ithaca, 1¢, June 14, 1940 (J. G. Franclemont) [CU]; 16, June 4, 1957; 10°, 19, June 14, 1957; 19, July 6, 1957; McLean Bogs Res., 1¢, July 4, 1924 [CU]; 29, July 22, 1924; 19, July 28, 1924; 10, July 6, 1946 (J. G. Franclemont) [CU]; 19, June 30, 1954 (J. G. Franclemont) [JGF]; 19, July 27, 1954; 1o¢, July 28, 1954; Varna, 16’, May 6, 1946 [CNC]; 1, June 17, 1946 [CNC]; 1o’, June 22, 1946 (E. G. Munroe) [CNC]; 14%, June 25, 1949 (Travassos and Rabella) [CNC]; West Chester Co., Katonah, 1o, July [ANS].
Norra Carouina: Polk Co., Tryon, 17, no date (Fiske) [USNM]; Robeson Co., Maxton, 1c’, May 23, 1944 (A. B. Klots) [AMNH]; Transylvania Co., Brevard, 19, June 13, 1942 (W. J. Westfall, Jr.) [CU]; 29, July 19, 1942; 2¢, July 23, 1942; 19, July 31, 1942.
Onto: Athens Co., Athens, 19, June 27, 1931 (W. C. Stehr) [UM]; Hamilton Co., Cincinnati, 1%, July 5, 1904 (Annette F. Braun) [ANS].
PENNSYLVANIA: Adams Co., Arendtsville, 1 @, 19, July 9, 1921 (S. W. Frost) [CU]; Allegheny Co., 2 2, July 1900 [CM]; 1 o, July 26 [CM]; Oak Station, 1 °, Aug. 2, 1908 (Fred Marloff) [CNHM];1 6, June 23, 1909; 1 o&, June 4, 1911 (Fred Marloff) [UK]; 1 &, June 14, 1911 (Fred Marloff) [CNHM];1 o, June 15, 1911 (Fred Marloff) [UK]; 1 9, June 24, 1911 (Fred Marloff) [CNHM]; Pittsburgh, 2 o', June 13, 1905 (Henry Engel) [CM];1 9, June 19, 1905 (Henry Engel) [CM]; 1 o', June 23, 1905 (Henry Engel) [USNM];1 9, July 9, 1905; 1 9, July 9, 1905 (Henry Engel) [CM]; 1 o, June 4, 1906 (Henry Engel) [CM]; 1 9, June 17, 1906; 1,1 9, June 18, 1906; 1 o, June 20, 1906; 1 9, June 26, 1906; 1 9, June 27, 1906; 1 o&, June 29, 1906; 1 &, July 1, 1906; 1 o&, July 6, 1906; 1 9, July 8, 1906 (Henry Engel) [USNM];1 o, July 10, 1906;1 9 [CM];207, July 15, 1906 (Henry Engel) [CM];1 9, Aug. 3, 1906; 2 9, July 15, 1907 (Hugo Kahl) [CM];1 &, July 16, 1907; 1 &, July 19, 1907; 1 &, June 23, 1908 (Henry Engel) [CM]; 1 9, June 24, 1908 (F. W. Friday) [LACM]; 1 ¢@, 1 2, June 27, 1909; 1 9, July 15, 1909 (Hugo Kahl) [CM]; 2 ¢, 1 9, July 8, 1910 (Henry Engel) [CM]; 1 &, June 13, 1911 (Hugo Kahl) [CM]; 1 9, June 15, 1911; 2 &, June 17, 1911; 1 9, Aug. 7, 1911; 1 @, July 12, 1912;2 @, July 2, 1914 [CM]; 1 &, July 22, 1914 [CM];1 @, July 24, 1916 (Henry Engel) [CM]; 1 ¢&, July 28, 1917 [CM]; 1 o, July 31, 1917 [CM];1 9, Aug. 17, 1917 [CM]; 1 &, June 6, (Henry Engel) [CM]; 1 &, June 17; 1 o', June 30; 1 o, July 3;3 9, July 11;1 9, July 12;1 ¢,1 9, July 14;1 9, July 15; 1 &, July 20; 1 o, July 24;1 #, 1 9, July 25; 1 9, August 2;1 @, Au-
34. U.S. NATIONAL MUSEUM BULLETIN 280
gust 3; Sharpsburg, 3 @, June 15-21 (Sweadner) [CM]; 2 o, June 22-30; 1 9, July 8-14; 6 3, 1 ?, July 15-21; 1 o@, August 15-21; Beaver Co., New Brighton, 1 9, July 2, 1902 (H. D. Merrick) [USNM];1 @, May 31, 1903; 1 ¢, July 11, 1904; 1 9, June 26, 1905 (H. D. Merrick) [UK]; 1 9, July 20, 1905 (H. D. Merrick) [USNM]; 1 o&, June 13, 1906 (H. D. Merrick) [UK]; 1 &, July13, 1906 (H. D. Merrick) [USNM]; Berks Co., Sinking Spring, 1 9 , June 29, 1952 (H. C. Moyer) [CNC]; Blair Co., Tyrone, 1 9, July 25, 1917 (J. G. Sanders) [BPI]; Bucks Co., Langhorne, 1 ?, June 21, 1924 (F. Haimbach) [ANS], Butler Co., Butler, 5 co, June 15, 1945 (Preston) [CM]; 4 o, June 17, 1945; 1 o, June 20, 1945; 2 o&, June 22, 1945; 1 o&, June 29, 1945; 7 o', 1 9, June 1945; 1 o, July 8, 1945; 1 0,1 9, July 15, 1945; 1 o, July 16, 1945; 2 o, July 17, 1945; 4 &, July 25, 1945; 5 o, 4 9, July 1945; 1 o, Aug. 7, 1945; 1 9, Aug. 17, 1945; 2 ot, POF August 1945; 1 #, 2 ?, July 1-7 (Sweadner) [CM]; Cumberland Co., Carlisle, 1 o’, June 25, 1931 (C. C. Hill) [USN M]; Dauphin Co., Rockville, 2 9 , July 22, 1917 [BPI]; 1 &, June 28, 1918 [BPI]; Fayette Co., Ohiopyle, 1 o, Aug. 10, 1905 [CM]; Forest Co., Kelletville, 1 #, July [CM]; Lycoming Co., Barbours, 1 o, July 8, 1921 (J. C. Bradley) [CU]; Monroe Co., Delaware Water Gap, 1 &, no date [AMNH]; Montgomery Co., Homebrook, Lower Merion Tp., 2 ?, June 23, 1916 [ANS]; 2 9, July 11, 1916 [ANS, CNC]; 1 9, July 18, 1916 [ANS]; 1 9°, July 19, 1916 [ANS]; 3 9, July 22, 1916 [ANS, CNC]; 2 92, July 23, 1916 [ANS]; 1 o, Aug. 2, 1916 [ANS]; 1 9, Aug. 3, 1916 [ANS]; 1 9, Aug. 7, 1916 [ANS]; 1 9, July 4, 1917 [ANS]; 1 9, July 8, 1917 [ANS]; 1 9, July 23, 1917 [ANS]; 1 9, Aug. 9, 1917 [ANS]; 1 2, June 10, 1920 [ANS]; 1 9, June 28, 1921 [ANS]; 1 ¢, 1 9, July 7, 1921 [ANS]; Spring Mount, 1 @, June 15 (H. A. Wenzel) [ANS]; Philadelphia Co., Philadelphia, 1 o&, June 16, 1909 (F. Weigand) [CNC]; 1 9, June 23 (F. Haimbach) [ANS]; Roxborough, 1 o&, June 22, 1913 [CNC]; 1 o&, June 24, 1913 [CNC]; 1 &, May 26, 1916 (F. Haimbach) [ANS]; Tioga Co., Blackwell, 1 ¢, July 1, 1921 (J. C. Bradley) [CU]; Washington Co., Finleyville, 2 @, June 8-14 [CM]; 1 9, June 24, [CM]; 1 o&, August 15-21 [CM]; 1 o, Sept. 22-30, [CM]; Westmoreland Co., Jeannette, 1 o, June 15 (H. G. Klages) [CM]; 1 6, June 19; 1 o, June; 16,3 9, July 8;2 9, July 19;1 6,19, July 25; 1 9, July 29:1 9; July 30;5 9, Aug. 6, 1904; 1 o, Sept. 19.
SoutH Carotina: Anderson Co., Anderson, 2, June 5, 1917 [CU].
Souta Daxora: Davison Co., Mitchell, 1 @, Sept. 14, 1945 (EK. C. Johnston) [CNC]; Union Co., Eik Point, 1 9, no date (C. N. Ainslie) [USNM]; county unknown, Cedar Pass, Bad Lands, 1 &, Aug. 13, 1940 (C. E. Mickel) [UM].
TENNESSEE: Davidson Co., Nashville, 1 @ ‘6/3’? (Ainslie) [USNM]; Grundy Co., Monteagle, 1 9, June 30, 1930 (Richards) [CU]; Knox Co., Knoxville, 1 9, June 28, 1916 (G. G. Ainslie) [USNM]; Morgan Co., Burrville, 1 #@, June 27, 1955 (A. K. Wyatt) [CNH M].
Texas: Culberson Co., Guadalupe Mts., 2 mi. north Pine Springs (5700 ft.), 1 9, July 19, 1963 (Harry Clench) [CM].
Uran: Tooele Co., Loop Camps, 13 mi. southwest Grantsville (7400 ft.), 1 &, July 19, 1958 (F., P., B., and J. Rindge) [AMNH]; 1 3, July 6, 1960.
Vermont: Rutland Co., Brandon, 1 ?, July 15, 1962 (J. C. Shaffer) [JCS].
Virainia: Giles Co., Mountain Lake, 1 o&, July 8, 1938 (L. J. and M. J. Milne) [CNC].
West Virainia: Greenbrier Co., White Sulphur Springs, 2 o, July 21, 1930 (J. G. Needham) [CU].
Wyomine: Carbon Co., Bottle Creek Camp. 7 mi. southwest Encampment (8800 ft.), 1 o&, Aug. 8, 1959 (F., P., and B. Rindge) [AMNH]; 1 o’, Aug. 9, 1959; 2 @, Aug. 11, 1959; Crook Co., 5 mi. west Alva, 3¢7, July 8, 1949 [AMNH; Reuter Canyon Camp, 5 mi. north Sundance (6100 ft.), 2 o, July 12, 1959
REVISION OF PEORIINAE AND ANERASTIINAE 35
(F., P., and B. Rindge) [AMNH]; 1 ao, July 13, 1959; 1 o&, July 14, 1959; (5900 ft.), 1 o&, July 11, 1962; Park Co., Cody, 5 &, no date [CM]; Lake Creek Camp, 13 mi. southwest Cooke City, Mont. (6900 ft.), 1 #@, July 25, 1959 (F., P., and B. Rindge) [AMNH].
CANADA: ALBERTA: Calgary, 1 o&, July 24, 1905 (F. H. Woolley-Dod) [CNC]; Horseshoe Canyon, Drumheller (2750 ft.), 1 #, July 17, 1960 (D. F. Hardwick) [CNC]; Lethbridge, 1 o&, July 30, 1921 (H. L. Seamans) [CNC]; 1 o&, June 27, 1956 (E. E. Sterns) [CNC]; 1 o&, July 7, 1956; Dominion Range Sta. Manyberries, 1 o, July 23, 1951 (D. F. Hardwick) [CNC]; 1 &, Aug. 2, 1951; 2 o, Aug. 3, 1951; 1 o, Aug. 4, 1941.
ManiToBa: Aweme, 1 oc’, June 23, 1904 (Criddle) [USNM]; 1 ¢, July 1, 1905: 3 o', July 2, 1921 (N. Criddle) [CNC]; 1 o, July 6, 1921; 1, 1922; 1 o, July 15, 1923; Brandon, 2 o&', 1 2, July 20, 1958 (R. B. Madge) [CNC]; 1 o&, July 31, 1958 (R. L. Hurley) [CNC]; Cartwright, 1 @, July 4, 1904 (E. F. Heath) [USNM]; 1 o’, July 26, 1905; 2 o&, July 29, 1905; 1 o&, no date; Red Rock Lake, Whiteshell For. R., 1 9, July 10, 1954 (C. D. Bird) [AMNH]; locality unknown, 1 ¢@, July 15 [USNM].
Nova Scotia: Annapolis, 1 o, June 20, 1946 (McDunnough and D. Ferguson) [CNC]; 1 o, July 5, 1946; 1 o&, Aug. 1, 1946; 1 &, Aug. 13, 1946; Petite Riviere, 20, July 21, 1935 (J. McDunnough) [CNC]; Truro, 19, July 9, 1913 (R. Mathe- son) [CU]; 1 o&, July 1, 1913; 1 9, Aug. 3, 1913.
Ontario: Blackburn, 1o’, June 18, 1941 (J. McDunnough) [CNC]; Black Sturgeon Lake, 1.7, July 23, 1963, [CNC]; Chatham, 19, June 27, 1931 (D. A. Arnott) [CNC]; 1¢, July 11, 1931 [CNC]; Constance Bay, 20’, July 9, 1934 (W. J. B.) [CNC]; Grand Bend, 10, July 17, 1939 (T. N. Freeman) [CNC]; Marmore, 1<’, June 13, 1952 (J. R. Vockeroth) [CNC]; 1, June 14, 1952; 3¢, 29, June 15, 1952; 16, June 17, 1952; 14, June 19, 1952;20¢, 39, June 20, 1952; 87,19, June 21, 1952; 1, June 22, 1952;1¢%, 29, July 3, 1952; 167, July 4,1952; 1¢, July 21, 1952; Mer Bleue, 1<’, June 12, 1937 (G. A. Hobbs) [CNC]; Mer Bleue, Hawthorne, 1¢’, June 15, 1937 (E. G. Lester) [CNC]; Normandale, 1<7, July 7, 1956 (Freeman and Lewis) [CNC]; Ottawa, 1¢@, June 30, 1907 (C. H. Young) [CNC]; Ottawa, Hunt Club, 27, July 8, 1937 (G. S. Walley) [CNC]; Palmer Rapids, 19, July 25, 1935 (F. A. Urquhart) [CNC]; Port Colborne, 1°, June 17, 1934 (J. J. de Gryse) [CNC]; 1, June 20, 1934; Sault Ste. Marie, 1¢, July 17, 1957 [CNC]; Simcoe, 27, June 9, 1939 (T. N. Freeman) [CNC, LACM]; 19,19, June 19, 1939; 167, June 20, 1939 (T. N. Freeman) [CNC]; 167, July 27, 1939; 10, June 5, 1949;19, July 5, 1949; 47, July 8, 1949; Strathroy, 27,19, July 5, 1926 (H. F. Hudson) [CNC, LACM]; 12’, July 11, 1929 (H. F. Hudson) [CNC]; Teeswater, 17, July 18, 1949 (D. F. Hardwick) [CNC]; 10, 19, July 20, 1949; 16, July 29, 1949; 2.7, Aug. 6, 1949; 19, Aug. 9, 1949; Toronto, 10%, June 1930 (H. 8S. Parish) [CU]; 10, July 1930; Trenton, 2, June 18, 1901 (Evans) [CNC]; 167, June 16, 1904; 19, June 28, 1907; 107, June 22, 1908; 1¢, June 14, 1911; 1.7, July 16, 1911; 106, June 27, 1912;19, July 6, 1912;1¢, July 12, 1912; Wanbamie, Parry Sound, 1, July 12, 1915 (H.S. Parish) [CU]; Wiarton 39, July 26, 1949 (D. F. Hardwick) [CNC].
QueseEc: Alcove, 19, July 4, 1935 (T. N. Freeman) [CNC]; Duchesnay, 1°, July 27, 1943 (R. Lambert) [CNC]; 16, July 14, 1947; Granby, 10’, June 22, 1938 (P. E. Mercier) [CNC]; Harrington Lake, Gatineau Park, 10, June 11, 1954 (J. E. H. Martin) [CNC]; 167, June 13, 1954 (H. J. Huckel) [CNC]; 107, June 15, 1954 (Martin) [CNC]; 29, June 16, 1954 (Huckel) [CNC]; 207, June 18, 1954 (Martin) [CNC]; 207, 29, June 20, 1954; 367, 19, June 21, 1954; 40, 29, June 24, 1954; 14, June 25, 1954; Knowlton, 14, July 7, 1929 (L. J. Milne) [CNC]; 19, July 11, 1929; Lac Mondor, Ste. Flore, 1¢, July 13, 1951 (E. G. Munroe)
36 U.S. NATIONAL MUSEUM BULLETIN 280
[CNC]; L. a la Torrue, 19, July 16, 1947 [CNC]; 106, 29, July 17, 1947 (E. G. Monroe) [CNC]; Meach Lake, 1, June 23, 1941 (G. A. Hobbs) [CNC]; 10, June 25, 1941 (G. A. Hobbs) [CNC]; Norway Bay, 19, July 9, 1937 (E. G. Lester) [CNC];19, July 15, 1937 (F. A. Urquhart) [CNC]; Ste. Anne de Bellevue, 1, June 29, 1947 (E. G. Monroe) [CNC]; 1¢, July 3, 1947; 1¢, 29, July 8, 1947; St. Annes, 26’, 39, July 5, 1947 (EH. G. Munroe) [CNC]; 10, 99, July 6, 1947:79, July 8, 1947;39, Aug. 2, 1947.
SASKATCHEWAN: Christopher Lake, 1o’, July 3, 1939 (A. R. Brooks) [CNC]; 1¢, July 5, 1939; Indian Head, 10, July 2, 1924 (J. J. deGryse) [CNC]; 1c, July 29, 1924; Saskatoon, 16’, June 30, 1923 (Kenneth M. King) [CNC]; 10, July 7, 1923; 16, July 17, 1923; 16, July 19, 1923; Willow Bunch, 10, July 27, 1955 (C. D. Miller) [CNC].
Discussion.—This is the most common and widely distributed member of the subfamily, the species being common from mid-June to early August throughout the northeastern United States. Western specimens tend to be larger and more uniform in maculation than those from the eastern and central states.
Peoria luteicostella (Ragonot), new combination
Figures 12, 82, 117, 155
Hypsotropa luteicostella Ragonot, 1887, p. 19; 1889, p. 117. — Hulst, 1890, p- 212, — Smith, 1891, p. 85. — Ragonot, 1901, p. 376. — Hulst, 1902, p. 489. — Barnes and McDunnough, 1917, p. 149.— Grossbeck, 1917, p. 134. — Hamp- son, 1918, p. 70. — McDunnough, 1939, p. 86. — Kimball, 1965, p. 251.
Wekiva nodosella Hulst, 1890, p. 215. — Smith, 1891, p. 85. — Hulst, 1902, p. 440. — Grossbeck, 1917, p. 134. — Rindge, 1955, p. 167.
Osceola perlepidellus Smith, 1891, p. 85. [Nomen nudum. New synonymy.]
Chipeta perlepidella Hulst, 1892, p. 62.— Barnes and McDunnough, 1917, p. 150. — McDunnough, 1939, p. 36. — Rindge, 1955, p. 169.— Kimball, 1965, p. 251. [New synonymy.]
Chipota [sic.] perlepidella Hulst, 1902, p. 441.
Bandera perlepidella (Hulst). — Hampson, 1918, p. 90.
Diacnosis.—This species resembles approximella in venation and maculation, but lacks the brownish white area posterior to the anal vein. The obliquely ascending palpi of both sexes and the postero- lateral gnathos hooks of the male are unique to this species.
DescriptTion.—Frons wine red; labial palpi obliquely ascending, wine red on outer sides, light pink on inner sides, third segment and dorsal half of second segment sometimes black on outer sides; maxillary palpi small, wine red; antennae with base wine red anteriorly, white posteriorly, shaft brown, wine red toward base, male laminate; outer sides of occiput, patagia, and tegulae wine red, vertex behind antennae and dorsal area of occiput white; legs white on outer sides, wine red on inner sides, tarsi black on inner sides.
Forewings with broad white band anterior to cell, narrowing to point at apex, bordered anteriorly near base with wine red to black, entire band sprinkled with wine red scales; white band bordered
REVISION OF PEORIINAE AND ANERASTIINAE 37
abruptly behind with black, black blending into posterior wine red area; outer fringe wine red to black. Hindwing light brown, darker in apical area. Venation as in P. bipartitella.
Male genitalia with medial process of uncus bearing minute central tooth, rarely several. Gnathos broadening posteriorly into a pair of large flat anteriorly directed hooks. Juxta V-shaped. Vinculum with dorsal margin flanged at apex. Valvae with costa terminating in strong slender spine reaching or surpassing the blunt membranous tip of cucculus; sacculus with numerous fine rather long hairs. Aedea- gus with vesica bearing small subserrate cornutus.
Female genitalia with ovipositor moderately setose laterally, numerous fine setae on caudal margin. Posterior apophyses tapering to point at base, not flattened, posterior half strongly curved; an- terior with base slightly thickened. Eighth abdominal segment unusually large.
Typrs.—H. luteicostella, in the Muséum National d’Histoire Naturelle; W. nodosella, in the U.S. National Museum (lectotype), in the American Museum of Natural History (lectoparatype); C. perlepidella, in the American Museum of Natural History.
Typr pata.—H. luteicostella, lectotype male, hereby designated, Florida; genitalia slide No. 535, J. Shaffer, June 20, 1965,
W. nodosella, lectotype male, hereby designated, Florida, Fernald collection; USNM 40081; genitalia slide No. 570, J. C. S., Nov. 23, 1965.
Lectoparatype: Florida; abdomen, antennae, and labial palpi lost.
In the original description of the species Hulst gives April as the date of capture.
C. perlepidella, lectotype female, hereby designated, Florida, collection G. D. Hulst; genitalia slide No. 3228, Carl Heinrich, June 14, 1946.
SPECIMENS EXAMINED.—28 o’, 29.
Distrisution (Map 10).—Southern Georgia and Florida.
UNITED STATES: Fuoripa; Alachua Co., Gainesville, 34, 19, July 8, 1927 (J. Speed Rogers) [CU]; 16, July 1927 (J. Speed Rogers) [CU]; Highlands Co., Archbold Biol. Sta., 14, June 17, 1964 (Jay C. Shaffer) [JCS]; 27, June 29, 1964; 307, July 5, 1964; 16, July 12, 1964; Hillsborough Co., Stemper, 14, Aug. 13, 1912 (G. Krautworm) [CM]; Volusia Co., Cassadaga, 1, Oct. 12, 1962 (S. V. Fuller) [CPK]; Wakulla Co., Panacea, 136’, 19, Aug. 11, 1926 (C. O. Handley) [USNM].
Grorata: County unknown, Billy’s Island, Okefenoke Swamp, 27, June 1912 [CU].
Anacostia, new genus
Type.—Anacostia tribulella, new species. Diacnosis.—The male genitalia are similar to those of Peoria, but differ mainly in the following ways: In Anacostia the spicate processes
38 U.S. NATIONAL MUSEUM BULLETIN 280
are three-branched rather than two-branched, the medial band of the uncus has a central pentagonal process, and the juxta bears a pair of lateral knobs, each with a small cluster of setae.
Description.—Labial palpi porrect, about 3% times as long as eye diameter; maxillary palpi moderate, not reaching frons; male antennae with basal segments of shaft fused, shaft serrate and fasciculate, female unknown; ocelli well developed.
Forewing with 11 veins; R, from cell relatively near to Rey; Re relatively long stalked with Rsis; Mo+3 stalked for about one-half their length or less; Cu, from near Mo+3;. Hindwing with 7 veins; Se and Rs stalked for about one-half their length; M; and Cu, stalked for about one-third their length.
Male genitalia with medial band of uncus very broadly rounded, bearing a well-developed, ventrally directed, apical, pentagonal process; spicate processes three-branched. Gnathos broad, flat, thin, broadened at base; sharp pointed apically, but without special process. Juxta V-shaped, bearing pair of setose knobs. Vinculum subquadrate, anterior corners well rounded. Valvae relatively slender; costa sparsely setose, with small apical tooth at tip of valve; sacculus densely pubescent. Aedeagus with vesica unarmed.
Female unknown.
Discussion.—The genus is apparently closely related to Peoria, but differences in the male genitalia are too great to allow inclusion of the one species in that genus.
Anacostia tribulella, new species FiaursEs 25, 83, 118
Dracnosis.—This is the only known species in the genus.
Description.—Labial palpi with all segments dark brown to black; antennae brown, shaft with tuft of black scales on basal seg- ments; head and thorax black; legs black on outer sides, brown on inner sides; abdomen dorsally brown, becoming golden brown anteriorly, ventrally black.
Forewing ground dark brown to black; area anterior to cell white sprinkled with black scales; discal spot, transverse anterior, transverse posterior, and subterminal lines black, not conspicuous against ground. Hindwing brown, darker toward apex; terminal line white, brown bordered on both sides.
Genitalia as described for the genus.
Typrs.—In the Academy of Natural Sciences (holotype); in the Cornell University collection (two paratypes).
Type pata.—Holotype, male, Holland Va. (Nansemond Co.), Aug. 1, 1945, O. Buchholz; A.N.S. Type No. 7817; genitalia slide No. 595, J. Shaffer, Apr. 24, 1966.
REVISION OF PEORIINAE AND ANERASTIINAE 39
Paratypes: Two males, both Clyde (Bryan Co.), Georgia, Sept. 11-12, 1931, Bradley and Knorr; C.U. Type No. 4404; one male genitalia slide No. 439, J. Shaffer, Mar. 25, 1965.
OTHER SPECIMENS EXAMINED.—None.
DistriBuTIon (Map 4).—Probably the Atlantic Coastal Plain; known only from Georgia and Virginia.
Arivaca, new genus
Typr.—Poujadia pimella Dyar, 1906.
Driaenosis.—Males of this genus may be recognized by the com- bination of (a) the gnathos bearing a bulbous or padlike medial process, and (b) each spicate process of the uncus having a short posterior and a much longer anterior sharp-pointed spine (figs. 119-125).
DescriptTion.—Frons conical, one-half to three-quarter times as long as eye diameter; labial palpi porrect; maxillary palpi very small; tongue poorly developed; antennae with scape compressed, male shaft with basal segments fused, female shaft filiform, scaled dorsally and laterally, finely ciliate ventrally, somewhat compressed; eye diameter about 0.2 mm greater in male than in female; ocelli normal.
Forewings with 10 or 11 veins; R, from well before upper outer angle of cell; Rs,4 stalked; M, from the angle; M;,3 stalked or fused, from lower outer angle; Cu, from just before the angle; Cu, from well before the angle. Hindwings with 6 or 7 veins; Sc and Rs approxi- mate or connate on basal half; M, from upper outer angle of cell; M, absent; Ms,Cu: free, stalked, or fused, from lower outer angle; Cu, from before the angle.
Male genitalia with spicate processes of uncus flat, each bearing short sharp posterior and long tapering anterior thorns. Gnathos arms straplike, bearing anteromedial projection; medial process bulbous or padlike. Vinculum broadly rounded. Aedeagus flattened, vesica bearing pair of cornuti.
Female genitalia with ductus bursae flattened, rather long, well sclerotized. Bursa small, lightly sclerotized. Ductus seminalis from posterior half of ductus bursae.
Discussion.—The seven species of Arivaca fall into four rather distinct subunits herein distinguished as species groups rather than as genera or subgenera. Because the genus is apparently best devel- loped in the unstudied region south of the United States, it is possible that intermediate forms exist between the species groups, and it is felt best not to give the groups nomenclatural status.
40 U.S. NATIONAL MUSEUM BULLETIN 280
Key to the Species Groups of Arivaca Based Upon the Maculation
i Cubitus with prominent white streak; white band anterior to cell (figs. 19, 2.) Mere h a See Re Cae See Utes Ie fare TN near teks ostreella group White trace of panies not Ce ae absent if white band present anterior GOCE), PF eReTe Bet A cL Pi. NCAR Sey fois AEN AML ys cee eaner eens 2 Mo+3 stalked; fee or secaias aipheer netenion to cell (figs. 16, 16). pimella group
Mo+3 fused; forewing somewhat lighter anterior to cell. . ...... 3 3. Forewing ground uniform reddish brown, prominent white band anterior foxcella@lorids) uae oe .. . . albicostella group Forewing gray, De eiaee of black and itty ponies radius and subcosta white traced (west of Mississippi River) .... . .. . albidella group
The Pimella Group
to
Driacenosis.—This group may be distinguished by the male valvae, in which the costa ends in a short triangular tooth well before the rounded tip of the cucullus. A similar projecting membranous cucullus exists in albicostella, but the costa terminates in a stout spine in that species.
Description.—Frons light brown to brown; labial palpi with basal segment white, second and third segments clothed with white-tipped brown scales dorsally and on outer sides, white ventrally and on inner sides; vertex light brown anteriorly, white posterior to antennae, occiput, patagia, and tegulae brown laterally, lighter dorsally; legs clothed with white-tipped brown scales on outer sides, white on inner sides.
Forewing ground light orange brown, veins white traced and bordered with brown, A; and A, traced for most of their length, A; traced at base; 11 veins; Mo; stalked. Hindwing light brown, fringe white; 7 veins; M; usually free from lower outer angle of cell.
Male genitalia with uncus moderately well sclerotized between spicate processes. Juxta rounded, with caudal V-shaped notch. Valvae with costa sparsely setose, apically bearing short triangular tooth; cucullus with rounded membranous tip projecting well beyond costa; sacculus with patch of stiff setae. Aedeagus with cornuti serrate, small, subequal.
Female genitalia with base of posterior apophyses flat, shaped like parallelogram. Bursa unarmed.
Discussion.—The two included species may be separated by the condition of vein R., which is stalked with R344 in linella but free in pimella, but this character is usually variable elsewhere in the sub- family so that positive identification should rest with examination of the male genitalia.
REVISION OF PEORIINAE AND ANERASTIINAE Al
Key to Species of the Pimella Group Based Upon the Male Genitalia
Posterior margin of medial process of uncus flat, usually bearing pair of short, sharp lateral teeth; apical process of gnathos padlike, not strongly projecting
(fig. 120) . Rein tite ene une typ as een nec dies ton ts a aaa Saas, a MeLLA Posterior eee of merit oe of uncus rounded, eeaeaeor apical process of gnathos bulbous, strongly projecting dorsally (fig. 119)... . . . . pimella
Arivaca pimella (Dyar), new combination
Fiaures 15, 50, 52, 84, 119, 156
Poujadia pimella Dyar, 1906, p. 31. — Barnes and McDunnough, 1917, p. 149. — McDunnough, 1939, p. 36.
Diacnosis.—The broadly rounded unarmed medial process of the uncus and the strongly projected medial process of the gnathos each separate this species from linella.
Description.—Antennae white, male shaft laminate, cilia about one-fifth as long as segment width.
Forewing sprinkled with brown and white anterior to cell; cell brownish white; Re free from cell. Hindwing light brown in female, somewhat darker in male.
Male genitalia with medial process of uncus rounded, unarmed. Gnathos with apical process strongly projected dorsally.
Type.—In the U.S. National Museum.
Type pata.—Lectotype male, hereby designated, Babaquivera Mts., Pima County, Ariz.; USNM 9012; genitalia slide No. 567, J. Shaffer, Nov. 23, 1965.
SPECIMENS EXAMINED.—45 a’, 369.
DisTRIBuUTION (Map 9).—Southern Arizona.
UNITED STATES: Arizona: Cochise Co., Cochise Stronghold, 1 ?, Aug. 30, 1958 (P. Opler) [UCB]; Paradise, 1 &#, 1 9, September (Poling) [CM]; Chiri- cahua, 1 mi. west portal, 1 @, 1 9, Sept. 10, 1950 (Cohn, Boone, Cazier) [AMNH]; Ramsay Canyon, Huachuca Mts., 1 @, 1 9, July 10-15, 1941 (A. B. Klots) [AMNH]; San Bernardino Ranch (3750 ft.), 1 9, August (F. Snow) [UK]; Southwestern Res. Sta., 5 mi. west Portal, 1 o, Sept. 6, 1959 (J. R. Bowers) [UCB]; 1 o, Aug. 25, 1962 (M. Statham) [AMNH];1 9, Sept. 2, 1959 (Carl W. Kirkwood) [CPK]; 1 9, Sept. 17, 1959; 1 9, Sept. 20, 1959; Gila Co., vicinity Globe, 1 &@, Aug. 4-5, 1937 (A. B. Klots) [AMNH]; Pima Co., ‘“Baboquavaria Mts.,” 4 o&, July 15-30, 1903 (O. C. Poling) [USNM]; 2 92, July 1-15, 1923; 2 @, July 15-30, 1923; 1 o, 29, Aug. 1-4, 1923; 1 9, Aug. 15-30, 1923;1 @, 1 9, Aug. 24-31, 1923; 47, 49, Sept. 1-15, 1923; 1 o, Oct. 1-15, 1923; 1 @, July 15-30, 1924; 7 &#, 1 9, Sept. 1-15, 1924; 1 o, Oct. 1-15, 1924; Brown Canyon, Baboquivari Mts., 1 @, Sept. 5, 1953 (Lloyd M. Martin) [LACM]; 1 o, Sept. 6, 1953; Madera Canyon, Santa Rita Mts., 1 9, Aug. 25, 1946 (J. A. Comstock, Lloyd M. Martin) [LACM]; 1 o&, Aug. 2, 1947; 39, Sept. 7, 1948 (Lloyd Martin) [LACM]; 1 o, Sept. 8, 1948; 1 9, Aug. 15, 1949 (C. W. Kirk- wood) [LACM]; 1 9, Aug. 17, 1949; 1 9, Sept. 11, 1950; 1 o&, 19, Sept. 13, 1950; 1 9, Sept. 15, 1950; 1 o, Sept. 5-12, 1951 (William Hammer) [CNC]; 1 9, Aug. 8, 1952 (C. W. Kirkwood) [LACM]; 1 o, Aug. 14, 1952; 2 9, Aug. 30,
42 U.S. NATIONAL MUSEUM BULLETIN 280
1952 (Lloyd Martin) [LACM]; 6 3,1 9, Aug. 19, 1953 (Robert J. Ford) [CNC]; 1 @, Aug. 16, 1953; 1 9, Aug. 18, 1953; 1 9, Aug. 21, 1953; 1 9, Sept..2, 1953 (Lloyd Martin) [LACM]; 1 2’, Sept. 8, 1953; 2 9, Aug. 24-Sept. 2, 1957 (W. A. Hammer) [LACM]; locality unknown: “So. Arizona,” 3 o, 1 9, August 1-15 (Poling) [USN M].
Discusston.—In the original description Dyar lists two males, one of these is in the National Museum, the fate of the other is unknown.
Arivaca linella, new species Ficures 16, 56, 85, 120, 157
Draenosis.—The flat, moderately well sclerotized medial process of the uncus distinguishes this species within the genus. The stalked condition of Re separates both sexes from pimella.
Description.—Antennae light brown, sublaminate in male, cilia about one-third as long as segment width, female with shaft white.
Forewings brown anterior to cell; Rz stalked with Rjy4.
Male genitalia with mediodistal margin of uncus irregular, usually bearing pair of small sharp teeth laterally. Gnathos with apical process padlike, not strongly projected dorsally.
Typrs.—In the Canadian National collection.
TyprE pata.—Holotype male, Colfax County, N. Mex., Sangre de Cristo Mts., Cimarron Canyon (7900 ft.), July 13, 1962, E. and I. Munroe; C.N.C. Type No. 9442; genitalia slide No. 274, J. Shaffer, Sept. 15, 1964.
Paratypes: Four males, same data and locality as holotype.
SPECIMENS EXAMINED.—18 6’, 39.
DistriBution (Map 9).
UNITED STATES: Arizona: Apache Co., 3 mi. west Eagar (7100 ft.), Pinon-Juniper life zone, 1 &, July 13, 1962 (E. and I. Munroe) [CNC]; Greer, White Mts. (8500 ft.), 1 #, Aug. 2, 1962 (KE. and I. Munroe) [CNC]; 1 9, Aug. 4, 1962; 5 o, Aug. 6, 1962; 1 9, Aug. 9, 1962; 1 o&, Aug. 11, 1962.
Cotorapo: ‘Rock Creek Canyon,” 10, July 19, 1961 (Margot May) [CPK].
New Mexico: Colfax Co., Cimarron Canyon, Sangre De Cristo Mts. (7900 ft.), 2 o&, July 14, 1962 (E. and I. Munroe) [CNC]; McKinley Co., McGaffey, Zuni Mts. (7500 ft.), 1 &, July 21, 1962 (EK. and I. Munroe) [CNC]; 1 <7, July 25, 1962; Sandoyal Co., Frijoles Canyon, Bandelier Nat. Mon. (6050 ft.), 4 #7, July 17, 1962 (E. and I. Munroe) [CNC]; 1 9, July 18, 1962; Horseshoe Springs Camp, 2 mi. west La Cueva (7900 ft.), 1 o&, July 29, 1961 (F., P., and J. Rindge) [ABK].
The Ostreella Group
Draanosis.—The group is easily recognized by the light orange forewings with the costal margin and cubitus both prominently marked with white. The tapering pointed valvae distinguish males of the group.
REVISION OF PEORIINAE AND ANERASTIINAE 43
DescripTion.—Frons about three-fourths as long as diameter of eye, orange laterally, lighter dorsally; labial palpi with basal segment white, second and third segments orange brown on outer sides, white on inner sides; antennae white, finely pubescent, bearing tuft of scales on basal segments of shaft; occiput and vertex white behind antennae, occiput brown laterally, patagia and tegulae light orange.
Forewing with broad white band anterior to cell, costa margined basally with dark brown; white band bordered posteriorly with narrow dark brown band dividing distally to trace Ry and M,; promi- nent white band tracing cubitus and Mo, broadest near lower outer angle of cell; Cu; and Cue traced with mixture of brown and white; Re from close to R3,4, sometimes short stalked. Hindwing with 7 veins; M; and Cu, from point or short stalked.
Male genitalia with medial process of uncus well sclerotized, rounded, bearing pair of caudomesally directed spines. Gnathos with medial process strongly projected. Valvae tapering to point.
Female genitalia with eighth abdominal segment heavily sclerotized, tapering posteriorly. Anterior apophyses strongly divergent; posterior parallel, pointed at base.
Key to the Species of the Ostreella Group Based Upon External
Characters M2+3 forked; white band extending from cell to costal margin. . . . ostreella M2+3 fused; white band extending half way from cell to costal margin, bordered Seperioniy.. Wath) Moh Orange, 0.2, <n cs ove) we oop An es sen poohella
Arivaca ostreella (Ragonot), new combination
Ficures 19, 53, 86, 121, 158
Saluria ostreeilla Ragonot, 1887, p. 18; 1889, p. 117. — Smith, 1891, p. 85.— Ragonot, 1901, p.362.—Hulst, 1902, p. 438. — Barnes and McDunnough, 1917, p. 149. — Hampson, 1918, p. 102. — McDunnough, 1939, p. 36.
Saluria ostrella [sic] Hulst, 1890, p. 211.
Peoria discostrigella Dyar, 1904, p. 115. [New synonymy.]
Diaenosis. — The stalked condition of Mo; distinguishes both sexes from poohella. Males are easily recognized by the pectinate antennae.
Description.—Labial palpi about 4% times as long as diameter of eye; male antennae pectinate.
Forewing with cell brownish orange; A; faintly traced on distal third; Az and A; basally traced with mixture of white and brown, Az usually broadly so; ground yellow or light orange posterior to cell; 11 veins; Me,3 stalked. Hindwing light brown in both sexes.
Male genitalia with uncus bearing heavy mediodorsal triangular plate, sharp pointed caudally. Juxta round, with deep V-shaped
285-934—68——4
44 U.S. NATIONAL MUSEUM BULLETIN 280
caudal incision. Valvae tapering to rounded tip; costa with small medial tooth. Aedeagus with vesica bearing one small serrate and one large pectinate cornutus.
Female genitalia with bursa unarmed.
Typrs.—S. ostreella, in the British Museum (Natural History) ; P. discostrigella, in the U.S. National Museum.
Type pata.—sS. ostreella, lectotype female, hereby designated, labeled as follows: ‘“Type; ARIZONA. Morrison.; Paravicine Coll. B.M. 1937-383.; Anerastia ostreella Rag. type; Saluria ostreella Ragonot; 2 genitalia slide I-4-1967 J. Shaffer No. 704.” P. discos- trigella, lectotype female, hereby designated, Roswell, New Mexico, August 22, Cockerell; USNM 7933; genitalia slide No. 566, J. C.5., Nov. 23, 1965. In the original description Dyar reports: “Three 99, Roswell, New Mexico, August 22 (T. D. A. Cockerell), Tucson, Arizona, July 21 (KE. A. Schwarz).”
SPECIMENS EXAMINED.—6 o’, 49 9.
Distrisution (Map 9).—Southern Arizona to Brownsville, Tex.
UNITED STATES: Arizona: Cochise Co., Southwestern Research Sta., 5 mi. west Portal (5400 ft.), 99, July 15, 1955 (W. J. Gertsch) [AMNH]; 19, July 26, 1955; 1 #, June 12, 1958; Pima Co., Redington, 3?, no date [USNM]; “Tuson,” 19, ‘21. 7.”? (E. A. Schwarz) [USNM].
New Mexico: Chaves Co., Roswell, 19, August (Cockerell) [USNM]; Eddy Co., White City, 39, May 14, 1950 (E. C. Johnston) [CNC]; 19, May 16, 1950; 1 #, 29, July 23, 1959, (A. B. Klots) [ABK]; Hidalgo Co., Lordsburg, 39 , Aug. 4, 1937 [AMNH]; Luna Co., 10 mi. east Deming, 3 o&, 29, July 12, 1917 [CU]; 149, July 8-15 [USNM].
Trxas: Brewster Co., Alpine, Davis Mts., 19 , Sept. 12, 1958 (R. R. McElvare) [CNHM]; Big Bend, 1 o&, Apr. 15-30, 1926 (O. C. Poling) [USNM]; 1 9, May 1-15, 1926; Cameron Co., San Benito, 29, July 8-15 [USNM];?, August [USNM]; Reves Co., 35 mi. northwest Pecos, 49, June 15, 1937 (George Willett) [LACM].
Arivaca poohella, new species Fiaures 20, 57, 87, 122, 159
Diacnosis.—The fused condition of M2,3 distinguishes this species from ostreella, as does the light orange trace on the costal margin.
Description.—Labial palpi 3% to 5% times as long as eye diameter; antennae sublaminate in male.
Forewing with white costal band extending half way from cell to costa, bordered anteriorly with light orange; cell orange; A; sometimes faintly traced near outer margin; A, and A; near base traced with white and bordered with dark brown; ground lighter orange posterior to A, fold; 10 veins; M,,; fused. Hingwing dark brown in male, light brown in female.
REVISION OF PEORIINAE AND ANERASTIINAE 45
Male genitalia with juxta rectangular; distally concave, proximally convex. Valvae tapering to point at apex of costa. Aedeagus with cornuti elliptical, serrate.
Female genitalia with bursa bearing near its base a cluster of about 20 stout, sharp-pointed, inwardly directed spines.
Types.—In the collection of John G. Franclemont, Cornell University.
TyprE pata.—Holotype male, Madera Canyon, 5600 ft., Santa Rita Mts., Santa Cruz County, Ariz., June 24, 1963, J. G. Franclemont; genitalia slide No. 714, J. Shaffer, Jan. 16, 1967.
Paratypes: Three males, two females, same data as holotype except as follows: two males, June 29, 1963, one with genitalia slide No. 715, J. Shaffer, Jan. 16, 1967; one male, June 28, 1963, genitalia slide No. 724, J. Shaffer, Jan. 27, 1967; one female, June 28, 1963, genitalia slide No. 716, J. Shaffer, Jan. 16, 1967.
OTHER SPECIMENS EXAMINED.—7 co’, 15 Q.
Disrrisution (Map 9).—Southern Arizona to Brownsville, Tex.
UNITED STATES: Arizona: Cochise Co., Southwestern Res. Sta., 5 mi. west Portal (5400 ft.), 1 9, May 15, 1956 (M. Statham) [AMNH];1 9, June 29, 1956 (Cazier and Ordway) [AMNH];1 9, July 5, 1956; 1 o&, July 9, 1956, 12, 1 9, July 11, 1958 (M. A. Cazier) [AMNH]; Pima Co., Baboquivari Mts., 10, July 1-15, 1923 (O. C. Poling) [USNM];1 ¢?, July 1-15, 1924;19, Aug. 15-30, 1924; Redington, 1 9, no date [USNM]; Santa Cruz Co., Madera Canyon, Santa Rita Mts. (5800 ft.), 1o7, 1 9, June 22, 1960 (David A. Wallesz) [CU]; 2 9, June 24, 1960; (4800 ft.) 1 9, June 18, 1963 (J. G. Franclemont) [J GF]; 2 9, June 29, 1963; 1 o, 1 92, July 1, 1963.
The Albidella Group
Diacnosis.—Males of the group are delimited by the broadly rounded apex of the valvae, the cucullus not exceeding the costa.
Derscription.—Frons of white-tipped brown scales; labial palpi about twice length of head, basal segment light brown, of white-tipped brown scales dorsally and on outer sides, white ventrally and on inner sides; antennae light brown, finely ciliate ventrally, tuft of scales on basal segments of shaft.
Forewing uniformly gray, a mixture of dark brown and white scales, Rs white traced; 10 veins; R2 stalked with R341; M2,; fused. Hindwing light brown; 7 veins, but M; sometimes completely fused with Cu.
Male genitalia with medial part of uncus poorly sclerotized, irregu- larly rounded. Apical process of gnathos well projected dorsally. Juxta subquadrate, anterior margin rounded, posterior with broad V-shaped notch. Valvae with costa setose, blunt tooth at apex and another just under half distance to base; tip rounded; sacculus finely setose. Aedeagus with cornuti equal or nearly so, serrate.
Female genitalia as in the Pimella Group.
46 U.S. NATIONAL MUSEUM BULLETIN 280
Key to the Species of the Albidella Group
Forewing appearing lighter posterior to A; fold; posterior cusp of spicate process a small triangular tooth about one-eighth length of anterior cusp (fig. 123)
albidella
Forewing not appearing lighter posterior to A; fold; posterior cusp of spicate
process a sharp thorn about one-fourth length of anterior cusp (fig. 124). artella
Arivaca albidella (Hulst), new combination
Ficures 17, 88, 123, 160
Peoria albidella Hulst, 1900, pp. 175, 439. — Barnes and McDunnough, 1917, p. 149.—McDunnough, 1939, p. 36.
Diacnosis.—Distinguishing features are given in the above key.
DerscrietTion.—Forewings light tan to grayish brown, appearing lighter posterior to A, fold.
Male genitalia with posterior cusp of spicate process poorly de- veloped, usually rounded, sometimes sharp pointed, usually no more than one-tenth length of anterior process.
Typre.—In the U.S. National Museum.
Type pata.—Lectotype male, hereby designated, Death Valley, “April 91 K.”; USNM 4809; genitalia slide No. 571, J. Shaffer, Nov. 23, 1965. In the original description Hulst notes: ‘“Death Valley, Cal.; taken by Mr. Koebele, in April. The type number of the National Museum is 4709.”
SPECIMENS EXAMINED.— 7 oc’, 1 9.
DistriButTion (Map 9).—Southwestern United States. UNITED STATES: New Mexico: Dona Ana Co., Mesquite, near Mesilla Park,? 2 #, July 12, 1917 [CU]. Texas: Reeves Co., Pecos, 4 7, 1 9, May 18, 1950 (E. C. Johnston) [CNC]; 1 &, June 2, 1950. Arivaca artella, new species
Fiaures 18, 89, 124, 161
Diaenosis.— Distinguishing features are given in the above key.
Derscription.—Male genitalia with posterior cusp of spicate process well developed as a sharp-pointed thorn about one-fourth as long as anterior cusp. Apical process of gnathos strongly projected, somewhat T-shaped in lateral view.
Typr.—In the Cornell University collection.
TypE pata.—Holotype male, Mesquite near Mesilla Park, N. Mex., July 12, 1917; Cornell University lot No. 542, sublot No. 46; lot No. 551, sublot No. 916; CU Type No. 4406; genitalia slide No. 366, J. Shaffer, Feb. 12, 1965.
2 Spelled Mesille Park on specimen labels.
REVISION OF PEORIINAE AND ANERASTIINAE 47
Paratypes: Three males, same date and locality as holotype, Cornell University lot No. 542, sublot No. 46.
OTHER SPECIMENS EXAMINED.—14 0,3 @.
DistRIBUTION (Map 9).
UNITED STATES: Arizona: Apache Co., St. Johns, 3 o&, July 26, 1937 (A. B. Klots) [AMNH].
New Mexico: Dona Ana Co., Mesquite near Mesilla Park, 10 o&, July 12, 1917 (from same lot as type series) [CU]; Hidalgo Co., Cienaga Ranch, near Rodeo, 1 9, July 12, 1948 (C. and P. Vaurie) [AMNH]; Sandoval Co., Jemez Springs (6200 ft.), 1 9, July 16, 1950 (Cohn, Boone, Cazier) [AMNH]; Sulphur Dam, 5 mi. north Jemez Springs (6300 ft.), 1 o&, July 17, 1950 (Cohn, Boone, Cazier) [AMNH]; county unknown, 1 9, no date [AMNH].
The Albicostella Group Arivaca albicostella (Grossbeck), new combination
Fiaurss 21, 60. 90, 125, 162
Calera (?) albicostella Grossbeck, 1917, pp. 134-135. — Klots, 1942, p. 419. Peoria albicostella (Grossbeck). — McDunnough, 1939, p. 36. — Kimball, 1965, p. 251.
D1aGnosis.—The presence of a white costal band coupled with the absence of white tracing on the cubitus distinguishes this species within the genus. In no other species of Arivaca does the male costa terminate in a subapical spine.
Description.—Frons brown; labial palpi about 2% times as long as eye diameter, basal segments light brown, second and third segments clothed with light brown tipped brown scales on outer sides, light brown on inner sides and ventrally; antennae with scape brown, shaft light brown, laminate, basal segments bearing tuft of scales, cilia slightly less than half as long as segment width; frons and vertex light brown dorsally, vertex brownish white posterior to antennae, occiput brownish white dorsally, brown laterally, patagia and tegulae brown; legs clothed with white tipped brown scales on outer sides, white on inner sides.
Forewing reddish brown, often sprinkled with light brown and dark brown scales; light brown anterior to cell, sometimes sprinkled rather heavily with darker scales. Hindwings light brown. Venation as in the Albidella Group.
Male genitalia with medial part of uncus weakly sclerotized, round- ed; spicate process with posterior thorn directed dorsally, then re- flexed 90° near base and caudally directed (fig. 125). Juxta rounded, posterior margin concave. Valvae with costa sparsely setose, produced as free spine before rounded membranous apex of cucullus; sacculus finely setose. Aedeagus with cornuti elliptical, serrate.
Female genitalia as in the Pimella Group.
48 U.S. NATIONAL MUSEUM BULLETIN 280
Typr.—tIn the American Museum of Natural History.
Type pata.—Lectotype female, designated by Klots (1942), Fort Myers, Fla., Apr. 23, 1912; genitalia slide No. 586, J. Shaffer, Nov. 28, 1965.
Lectoparatype female, Everglades, Fla., Apr. 7, 1912.
SPECIMENS EXAMINED.—10 o’, 3 @Q.
DistrRiIBuTION (Map 10). — Known only from southern Florida.
UNITED STATES: Fior1pa; Broward Co., Ft. Lauderdale, 9 , Mar. 27, 1928 (D. M. Bates) [CPK]; Collier Co., Everglades, 4 o&, April 1-7 [USNM]; 3 ¢, April 8-15 [USNM]; Dade Co., Paradise Key, Everglades National Park, 9, Apr. 2, 1952 (G. S. Walley) [CNC]; o, Apr. 3, 1952; Sarasota Co., 9, May 13, 1946 (C. P. Kimball) [CPK]; Siesta Key, @, Mar. 31, 1952 (C. P. Kimball) [CPK]; 3, Mar. 30, 1954 (C. P. Kimball) [CPK].
Atascosa Hulst
Atascosa Hulst, 1890, p. 210. — Smith, 1891, p. 85. — Hulst, 1902, p. 438. [Type: Atascosa bicolorella Hulst, 1890. Original designation.]
Draenosis. — The flat, elbowed, bluntly pointed spicate processes of the uncus (fig. 126) are diagnostic of the genus.
Description. — Frons conical; labial palpi upturned with third segment decumbent in males, porrect in females; maxillary palpi reaching frons, cylindrical tuft of long slender scales, sometimes ex- panded and fanlike; antennae sublaminate, ciliate in males, filiform, finely ciliate in females; ocelli well developed.
Forewing with 11 veins; R, free from cell; R, from near to or stalked with Rsi4; M2,3 stalked for about half its length. Hindwing with 7 veins; Sc and Rs stalked; M, from upper outer angle of cell; M, absent; M3 stalked with Cu,, from lower outer angle; Cu, from just before the angle.
Male genitalia with spicate processes in form of a pair of flat lateral arms, ventroposteriorly directed from base, then elbowed and distal half ventrally directed; arms bluntly pointed. Gnathos with apical process a small posteroventrally directed thorn; arms straplike. Juxta subquadrate, shieldlike. Vinculum \V-shaped, somewhat flattened anteriorly. Valvae with costa bearing subapical spine; sacculus with numerous stout setae; cucculus rounded, membranous, surpassing costa. Aedeagus flattened, vesica unarmed.
Female genitalia with ovipositor moderately setose, caudal margin finely setose. Apophyses fairly straight, of about equal length; well developed; base of posterior shaped like a parallelogram; anterior curved downward near base. Bursa unarmed. Ductus seminalis from posterior end of bursa.
REVISION OF PEORIINAE AND ANERASTIINAE 49
Atascosa glareosella (Zeller) FiauReEs 26, 54, 91, 126, 163
Anerastia glareosella Zeller, 1872, pp. 553-554.
Saluria glareosella (Zeller). — Ragonot, 1889, p. 117. — Hulst, 1890, pp. 211- 212. — Smith, 1891, p. 85. — Hampson, 1918, pp. 95-96.
Atascosa bicolorella Hulst, 1890, pp. 210, 227. — Smith, 1891, p. 85. — Rindge, 1955, p. 158.
Maricopa albocostella Hulst, 1900, p. 176 [described in Phycitinae]; 1902, p. 438. — Rindge, 1955, p. 157. — Heinrich, 1956, p. 316 [transferred to Aner- astiinae]. [New synonymy.]
Poujadia glareosella (Zeller). — Ragonot, 1901, pp. 345-346. — Barnes and McDunnough, 1917, p. 149. — Forbes, 1923, p. 638. — MceDunnough, 1939, p. 36.
Atascosa glareosella (Zeller). — Hulst, 1902, p. 488. Valdiva albocostella (Hulst). — Barnes and McDunnough, 1917, p. 148. — Mce- Dunnough, 1939, p. 34.
Driacnosis. — This is the only known species in the genus.
Description.—Frons dark brown, with red posterodorsal patch; labial palpi with basal segments white, second and third segments dark brown on outer sides, ventrally white sprinkled with red; anten- nae with scapes white on outer sides, mixture of red and brown on inner sides, shaft light brown; vertex and occiput brown, lighter dorsally; patagia, tegulae, and dorsum of thorax dark brown, usually sprinkled with red.
Forewing with white costal band extending posterior to middle of cell, narrowing to point at apex, sprinkled with grayish red, grayish- red basal dash; white band bordered posteriorly by dark brown band, mixture of grayish red and dark brown distal to cell; posterior of grayish red and dark brown distal to cell; posterior third of wing grayish red sprinkled with brown; transverse posterior band dark brown; discocellular traced with dark brown, forming discal spot; fringe brown, with three white lines. Hindwing light brown, darker toward apex.
Genitalia as described for the genus.
Typrs.—A. glareosella, in the British Museum (Natural History), (from Zeller collection); A. bicolorella and M. albocostella, in the American Museum of Natural History.
Typp pata.—a. glareosella, lectotype female, hereby designated, labeled as follows: ‘Type; 15/8; Bosque Co. Texas; Zell. Coll. 1884.; Anerastia glareosella Z. N.A.I. 553 [green label in Zeller’s handwrit- ing]; 2 genitalia slide I-4-1967 J. Shaffer No. 709.”
A. bicolorella, lectotype male, hereby designated, Blanco County, central Texas, August, collection G. D. Hulst; genitalia slide No. 3222, Carl Heinrich, June 14, 1947.
50 U.S. NATIONAL MUSEUM BULLETIN 280
M. albocostella, lectotype male, hereby designated, Anglesea, N.J., August 21?, collection G. D. Hulst, genitalia slide No. 3219, Carl Heinrich, June 14, 1946. Lectoparatype male, Anglesea, N.J., August 21?, collection G. D. Hulst.
SPECIMENS EXAMINED.—44 o’, 18 Q.
DistriBuTION (Map 12).—Atlantic and Gulf Coastal Plain, Texas to Florida, north to Massachusetts.
UNITED STATES: Cotorapo: El Paso Co., Colorado Springs, Fountain Valley School, 1 o&, July 7, 1935 [AMNH] (see discussion section).
Connecticut: New Haven Co., East River, 1 o&, Aug. 12, 1907 (Charles R. Ely) [USNM]; 1 o, Aug. 16, 1907; 2 &, July 19, 1908; 27, July 20, 1908; 1 o, July 22, 1908; 1 o&, July 24, 1908; 2 o, 1 9, July 27, 1908; 1 o, July 28, 1908; 1 o', July 30, 1908; 1 o, Aug. 1, 1908; 1 9, Aug. 2, 1908; 1 9., Aug: 7, 1909; 1 o', Aug. 10, 1909; 1 o&, Aug. 12, 1909;1 &, 1 9, July 1910; 1 9, July 19, 1912; 1 o, Aug. 1, 190?.
Fioripa: Dade Co., Homestead, 1 o&, Aug. 7, 1963 (D. O. Wolfenbarger) [CPK].
MassacuusEtts: Barnstable Co., Barnstable, 1 #7, July 4, 1949 (C. P. Kimball) [CU];1 o&, July 10, 1949 (C. P. Kimball) [CU]; 2 #, July 10, 1949 (C. P. Kimball) [CPK]; 1 9, July 12, 1949; 1 o@, Aug. 1, 1952; West Barnstable, 3 o, July 15, 1949 (C. P. Kimball) [CPK]; 1 ¢&, July 16, 1949 (C. P. Kimball) [CNC]; 2 ¢ 4 9, [CU]; 1 o, July 17, 1949 (C. P. Kimball) [CU]; 1 @, July 18, 1949 (C. P. Kimball) [CPK];1 9, July 20, 1949; Dukes Co., Martha’s Vineyard, 1 o, Aug. 11, 1946 (F. M. Jones) [CNC]; 1 o&, July 18 (F. M. Jones) [CPK].
New Jersey: Bergen Co., Oakland, 1 <&, July 26, 1947 (C. P. Kimball) [CPK]; 1 6, July 27, 1947 (C. P. Kimball) [CU]; 1 9, Aug. 4, 1948 (C. P. Kimball) [CPK]; 1 &, Aug. 9, 1948 (C. P. Kimball) [CU]; 1 o&, Aug. 13, 1948; 1 o&, Aug. 14, 1948; 1 @, Aug. 24, 1948; Burlington Co., New Lisbon, 1 o&, July 31, 1931 (E. P. Darlington) [ANS]; 1 9, July 14, 1933; 1 @, Aug. 10, 1939; 1 9, July 31, 1942; 1 o&, Aug. 4, 1942; Morris Co., Mendham, 1 o&, Aug. 3, 1935 (G. H. Tate) [AMNH]; Ocean Co., Lakehurst, Wrangle Brook Road, 1 9, Aug. 30, 1956 (J. G. Franclemont) [J GF].
New York: Suffolk Co., Riverhead, 1 o&, July 1, 1933 (Roy Latham) [CU].
Nortu Carouina: Polk Co., Tryon, 1 9, Aug. 11, 1904 (Fiske) [USNM]; 1 9, Aug. 14, 1904.
Discussion.—The Colorado specimen has wing venation and male genitalia typical of glareosella, but differs in having uniform brownish- orange forewings, unmarked except by a faint discal spot, and in being slightly larger than typical members of the species. The specimen may or may not be glareosella.
Homosassa Hulst
Homosassa Hulst, 1890, p. 214. — Smith, 1891, p. 85. — Haulst, 1902, p. 440. [Type: Ephestia ella Hulst, 1887. Monobasic.]
Draenosis.—The long, straight, unbranched ventrolaterally di- rected spicate processes of the uncus are diagnostic of the genus.
REVISION OF PEORIINAE AND ANERASTIINAE 51
Description.—Labial palpi of males obliquely ascending with third segment somewhat decumbent, two to 2% times as long as eye diameter, females porrect, three times eye diameter; maxillary palpi small; tongue rudimentary; antennae sublaminate and ciliate in males, filiform and finely ciliate in females; ocelli normally developed.
Forewing with 10 or 11 veins; R, free from cell or stalked with R344; Mo+3 stalked or fused, from lower outer angle of cell; Cu, from just before the angle. Hindwing with 7 veins; Sc and Rs stalked; M, from upper outer angle; M, lost; M; stalked with Cu,, from lower outer angle of cell; Cu, from just before the angle.
Male genitalia with spicate processes of uncus straight, unbranched, ventrolaterally directed, tapering to broad triangular base.
Juxta shield-shaped. Valvae subrectangular with costa projecting, bluntly rounded and flattened apically; sacculus rather densely setose. Aedeagus with thickened ring on posterior end.
Female genitalia with ovipositor strongly compressed. Apophyses rather straight, about equal in length. Bursa unarmed. Ductus seminalis from posterior end of bursa.
Key to the Species of Homosassa
freee ere tSedn(He Oye alae sk fie as es ee PS eo, he Se ew eee, owe ella Mis stalked) (fig. S54) Gy.) oto, Avi beaty, Ria tt Ce 2. Male with uncus bearing well sclerotized ring Ge 129) Xoo) >t eincudela Male with uncus not bearing sclerotized ring (fig. 128). . . .. . . platella
Homosassa ella Hulst
Fiacures 27, 58, 92, 127, 164
Ephestia ella Hulst, 1887. — Rindge, 1955, p. 161. Anerastia ella (Hulst). — Ragonot, 1889, p. 117; 1901, p. 400. — Barnes and McDunnough, 1917, p. 149. — McDunnough, 1939, p. 36. — Kimball, 1965, Seok, Perse ella Hulst, 1890, pp. 214, 227. — Smith, 1891, p. 85. — Hulst, 1902 p. 440. — Grossbeck, 1917, p. 134. Rhinaphe ella (Hulst). — Hampson, 1918, p. 86.
Diacnosis.—The loss of vein M, distinguishes this species from the other two members of the genus.
Descrirtion.—Frons conical, dark brown; labial palpi with basal segments white, second and third segments dark brown dorsally, white ventrally and on inner sides; antennae with scape dark brown anteriorly, white posteriorly, shaft light brown; vertex, occiput, patagia, tegulae, and dorsum of thorax brown to dark brown; legs brown, lighter on inner sides, tarsal scales white tipped.
Forewing with white band anterior to cell, tapering to point at apex; proximal half of band margined anteriorly with dark brown line broadened at base; white band usually sprinkled with red; ground
52 U.S. NATIONAL MUSEUM BULLETIN 280
uniform dark brown posterior to white band, sometimes with reddish hue posterior to A; fold; 10 veins; R, from well before upper outer angle of cell; R, free from cell or stalked with R3,4; M2,3 fused. Hind- wing brown or light brown, darker toward apex.
Male genitalia with uncus bearing well sclerotized ring, open anteriorly and forming pair of ventrally directed sharp-pointed spines. Gnathos (fig. 127) pi-shaped, transverse portion upturned and sharp, pointed laterally. Aedeagus with vesica bearing single sharply serrate inconspicuous cornutus.
Female genitalia as described for the genus.
Typrs.—In the American Museum of Natural History (lectotype) ; in the U.S. National Museum (lectoparatype).
TYPE DATA.—Lectotype male, hereby designated, Florida, collection G. D. Hulst; genitalia slide No. 3225, Carl Heinrich, June 14, 1946.
Lectoparatype: Male, Fernald collection, USNM 40079; genitalia slide No. 572, J. Shaffer, Nov. 23, 1965.
In the original description Hulst lists three males and two females from Florida.
SPECIMENS EXAMINED.—90 go’, 42 9.
DistriBuTion (Map 11).—Atlantic and Gulf Coastal Plain, eastern Texas to Florida, north to North Carolina.
UNITED STATES: AtaBama: Mobile Co., Theodore, 1 o, June 12, 1917, [CU].
Froriwa: Alachua Co., Gainesville, 1 @, 29, July 8, 1927 (J. Speed Rogers) [CU]; 1 &, July 10, 1927; 1 &, Apr. 22, 1952 (O. Peck) [CNC]; Charlotte Co., Punta Gorda, 2 ¢, Apr. 12, 1952 (G. 8. Walley) [CNC]; Dade Co., Florida City, 2 o, Apr. 17, 1947 (Otto Buchholz) [ANS]; 1 o, Apr. 20, 1947; Homestead, 1 o&, Feb. 24,1959 (D. O. Wolfenbarger) [CPK]; 1 o, Apr. 15, 1959;1 o, Apr. 16, 1959; 2 &, July 17, 1959; 1 of, Feb. 22, 1955; 1 &, July 29, 1958; Royal Palm Hammock (also called Paradise Key, or Royal Palm State Park), Everglades National Park,1<, Apr. 11 (F. M. Jones) [ANS]; 1 o, 1930; 1 o&, Mar. 15, 1938 (E. P. Darlington) [ANS]; 1 o, Mar. 17, 1938; 1 &, Mar. 20, 1938; 2 0, 2 9, Apr. 2, 1952 (G. S. Walley) [CNC]; 1 9, Apr. 3, 1952; Flager Co., Pellicer Crk., 13 mi. north Bunnell, 1 9, May 8, 1954 (J. Bauer) [CM]; 1 o, Apr. 10, 1954; 1 9, Apr. 24, 1954; Highlands Co., Archbold Biol. Sta., 1 9, Apr. 27, 1947 (J. G. Needham) [CU]; 1 o&, Apr. 28, 1947; 2 9, May 14, 1947; 2 9, June 17, 1964 (Jay C. Shaffer) [JCS]; 1 &%, 3 2, June 18, 1964;1 0,1 2, June 20, 1964; 2 o, 1 9, June 22, 1964; 2 &, June 25, 1964; 2 #, 1 9, June 26, 1964; 2 ¢, June 28, 1964; 1 9, June 29, 1964; 1 9, July 2, 1964; 1 o, July 3, 1964; 1 o, July 5, 1964; 1 o&, July 11, 1964; 1 9, July 12, 1964; 1 o&, July 13, 1964; Hillsborough Co., Stemper, 1 o&, Aug. 19, 1912 (G. Krautwurm) [CM]; 2 2, 1 9, Sept. 1, 1912; 2 &, Sept. 4, 1912; 2 #, Sept. 5, 1912; 2 o, Sept. 6, 1912; 1 o, Sept. 9, 1912;1 #, 1 9, Sept. 18, 1912; 1 9, Oct. 1, 1912; 1 o&, Oct. 8, 1912; 1 &, Oct. 9, 1912; 1 &, June 4 [CM]; 1 9, June 19 [CM]; Lake Co., Leesburg, 1 o&, May 14, 1961 (C. H. Curran) [AMNH]; Manatee Co., Gulf Coast Exp. Sta., Bradenton, 1 o’, May 11, 1955 (E. G. Keisheimer) [CPK]; Oneco, 1 ?, May 5, 1953 (Paula Dillman) [CPK]; 1 &, May 8, 1953; 1 9, May 19, 1953; 1 o', May 25, 1953; 1 9, June 6, 1953; 1 o&, 1 9, June 7, 1953; 1 9, June 10,
REVISION OF PEORIINAE AND ANERASTIINAE 53
1953; 1 o, July 15, 1953; 1 o&, Oct. 15, 1953; 1 o&, June 1, 1954; 1 9, June 9, 1954; Pinellas Co., Dunedin, 1 o&, Apr. 22, 1925 (W. 8. Blatchley) [ANS]; 2 ¢, May 24, 1925; Sarasota Co., 1 o&, May 7, 1946 (C. P. Kimball) [CPK]; 1 9, May 12, 1946;1 o&, May 19, 1946; Siesta Key, 1 9, May 21, 1956 (C. P. Kimball) {[CPK]; St. Johns Co., Hastings, 6 @, 2 9, April [USNM]; 5 o, 1 9, May [USNM]; 3 3#, 1 2, June [USNM]; 1 o, Sept. 30 [USNM]; county unknown, “Altmont,’” 1 9, Sept. 21, 1924 (F. R. Colo) [CU]; locality unknown, 1 a, no date [USNM].
Groraia: Charlton Co., 1 o&, June 7, 1946 (Otto Buchholz) [ANS 16, June 9, 1946; 27, June 10, 1946; 1 o&, June 11, 1946.
Mississippi: Jackson Co., Biloxi, 1 @, June 13, 1917 [CU].
Nort Carouina: Brunswick Co., Leland, 1 ?, June 17, 1946 (Otto Buchholz) [ANS]; 2 o&, June 20, 1946; 1 9, Sept. 2, 1946; Robeson Co., Maxton, 1 ¢, May 1-15, 1943 (A. B. Klots) [AMNH].
Texas: Burnet Co., 1 o&, no date (F. G. Schaupp) [USNM]; Nueces Co. Corpus Christie, 1 @, Mar. 26, 1943 (W. M. Gordon) [CU].
Discusston.—In many specimens the distal one-half of each spine of the uncus ring is bent anteroventrally (fig. 127), and the anteriorly directed spine of the gnathos is posterior (fig. 127a) in relation to that of the type (fig. 1276). Specimens of both types have been collected at the Archbold Biological Station and do not represent geographical varieties. In view of the small differences between the two types, they are held to be conspecific.
Homosassa platella, new species FicureEs 28, 93, 128
Diacenosis.—The broad flat transtilla and lack of a sclerotized ring on the uncus each will distinguish this species.
Description.—Maculation similar to that of ella. Forewing with 11 veins; Mo,; stalked.
Male genitalia with uncus lacking sclerotized ring.
Gnathos expanded into pair of large, flat, medially fused, posteriorly bicuspitate or tricuspitate plates. Juxta four-sided with medial \- shaped heavy sclerotization. Aedeagus with vesica bearing pair of weakly sclerotized sharply serrate cornuti.
Female unknown.
Typrs.—In the U.S. National Museum.
Typr pata.—Holotype, male, Hastings, Fla. (Saint Johns Co.), March, collection W. D. Kearfott; USNM 69385; genitalia slide No. 596, J. Shaffer, Apr. 26, 1966.
Paratypes: Three males, Hastings, Fla., collection W. D. Kearfott; two dated March, one male genitalia slide No. 103, J. C. 8., Dec. 30, 1963; one dated April, male genitalia slide No. 493, J. Shaffer, Apr. 13, 1965.
OTHER SPECIMENS EXAMINED.—4 oJ.
Distrisution (Map 11).—Known only from Florida.
54 U.S. NATIONAL MUSEUM BULLETIN 280
UNITED STATES: Ftortpa: Collier Co., Everglades, 1 o&, Apr. 6, 1912 [AMNH]; Dade Co., Homestead, 1 o, Apr. 28, 1952 (J. R. Vockeroth) [CNC]; Pinellas Co., Dunedin, 1 o, Feb. 23, 1921 (W. 8S. Blatchley) [ANS]; Saint Johns Co., Hastings, 1 #, May [AMNH].
Homosassa incudella, new species Ficgures 29, 94, 129, 165
Diaenosis.—The sclerotized ring of the uncus bears a pair of two- pointed processes rather than a simple process as in ella, or with the ring absent as in platella. Female specimens may be distinguished from those of ella by the presence of a dorsal invagination on the eighth abdominal segment.
Description.—Maculation similar to that of ella. Forewing with white costal band sewn with dark brown scales; 11 veins; Me,; stalked.
Male genitalia with uncus similar to that of ella, but anterior processes of ring each anvil-shaped with sharp-pointed anterior and posterior processes. Gnathos with pair of subapical thickened proc- esses, each bearing anterior and posterior spines. Aedeagus with vesica unarmed.
Female genitalia with dorsal invagination on eighth abdominal segment, and smaller invagination dorsally on the membrane between segments eight and nine.
Typrs.—In the Canadian National collection.
Typr pata.—Holotype, male, Lake Murray, Love and Carter Counties, Oklahoma, May 20, 1950, W. J. Reinthal, ‘339/50’; C.N.C. Type No. 9441; genitalia slide No. 546, J. Shaffer, July 15, 1965. A mass of glue supports the specimen on the pin from beneath, and I have added a small amount of glue beneath the right wings to support them.
Paratype: Female, data as given for the holotype; genitalia slide No. 548, J. Shaffer, July 16, 1965.
OTHER SPECIMENS EXAMINED.—None.
DisTRIBUTION (Map 11).
Reynosa, new genus
Typrn.—Atascosa floscella Hulst, 1890.
Diagnosis.—The short, stout, sharp-pointed spicate processes of the uncus are diagnostic of the genus.
DescriptTion.—Frons conical; labial palpi porrect, about twice as long as eye diameter in males, about 2% times in females; maxillary palpi reaching frons or nearly so, spreading; tongue not exposed between palpi; antennae filiform to subserrate in both sexes, com- pressed, ciliate ventrally, male shaft with two basal segments fused; ocelli present.
REVISION OF PEORIINAE AND ANERASTIINAE 55
Forewing with 11 veins; R, usually free from cell, sometimes stalked with R3,4; M; from upper outer angle of cell; M.,3 stalked about one- half their length, from lower outer angle; Cu, from just before the angle. Hindwing with 7 veins; Sc and Rs well stalked; M, from upper outer angle of cell; M; and Cu, long stalked, from lower outer angle; Cur from cell very near to M; and Cu.
Male genitalia with uncus bearing pair of short apical lobes; spicate processes a pair of short, stout, sharp-pointed anteriorly directed, ventral hooks. Gnathos bearing small, sharp-pointed, posteriorly directed apical and pair of subapical hooks. Juxta subquadrate, an- terior margin convex, posterior concave. Vinculum well developed, rounded. Valvae with cucullus membranous and projecting beyond costa; sacculus rather densely pilose. Aedeagus untapered, vesica unarmed.
Female genitalia with ovipositor compressed. Apophyses rather straight, anterior and posterior about equal in length. Ductus bursae short. Bursa unarmed, with long slender neck. Ductus seminalis from posterior end of bursa, slender.
Reynosa floscella (Hulst), new combination FicuREs 22, 95, 130, 166.
Atascosa floscella Hulst, 1890, pp. 210-211. — Smith, 1891, p. 85. — Hulst, 1902 p. 438. — Rindge, 1955, p. 162.
Poujadia floscella (Hulst). — Ragonot, 1901, p. 346, — Barnes and MecDunnough 1917, p. 149. — McDunnough, 1939, p. 36.
Saluria floscella (Hulst). — Hampson, 1918, p. 96.
Diaenosis.—This is the only known species in the genus.
Description.—Labial palpi with basal segments white, second and third segments brown to reddish brown on outer sides, white on inner sides; frons, occiput, patagia, and tegulae brown.
Forewing with ground light brown; costal area and cell white, sprinkled with reddish-brown scales; prominent dark brown orbicular spot and line extending posteriorly from spot to inner margin forming second spot on A, fold; dark brown transverse posterior beginning with prominent spot between M, and Cu, continuing to inner margin; terminal line of dark brown dots. Hindwing light brown on both sides.
Genitalia as described for the genus.
Typrs.—In the U.S. National Museum (lectotype) ; in the American Museum of Natural History (lectoparatype).
TypsE pata.—Lectotype female, hereby designated, Blanco County, central Texas; Fernald collection; USNM 40078; genitalia slide No. 594, J. Shaffer, Apr. 23, 1966.
Lectoparatype: Male?, Blanco County, central Texas, collection G. D. Hulst; abdomen lost.
?
56 U.S. NATIONAL MUSEUM BULLETIN 280
SPECIMENS EXAMINED.—10 go’, 12 9.
Disrrinution (Map 12).—Known only from Gulf Coastal Plain of Texas.
UNITED STATES: Texas: Cameron Co., Brownsville, 1 9, May 4, 1904 (H. S. Barber) [USNM]; 1 92, no date [USNM]; 1 o&, Mar. 19, 1937 (T. N. Freeman) [CNC]; 2 o&, Mar. 22, 1937; San Benito, 1 9, June 16-23 [USNM]; 1 9, July 8-15 [USNM]; 1 9, July 24-31 [USNM]; 1 9, August [USNM]; 1 9, Aug. 1-7 [USNM];1 o&,1 9, Sept. 8-15 [USNM]; 1 &, Mar 16-23 [USNM]; Fort Bend Co., Richmond, Brazos River, 1 @, June 22, 1917 [CU]; Nueces Co., Corpus Christie, 1 o&, 2 9, Apr. 15, 1943 (W. M. Gordon) [CU]; 1 2 May 22, 1943; 1 9, June 4, 1943; 1 o&, June 20, 1943; 1 o&, 1 9, Sept. 25-Oct. 15, 1943.
Goya Ragonot
Goya Ragonot 1888, p. 43. [Type: Goya albivenella Ragonot, 1888. Designated by Hampson in Ragonot 1901, p. 349.]
Driaenosis.—The unusual male genitalia make this genus quite distinctive within the subfamily; the presence of a transtilla and lateral subrhomboid processes of the uncus are both unique to Goya.
Dezscription.—Labial palpi about twice as long as eye diameter; tongue rudimentary; antennae with basal segments fused in male; eye large; ocelli present.
Forewing with 11 veins; R» free from cell near R344 or stalked with the latter; M, from upper outer angle of cell; M2,3; stalked, from lower outer angle. Hindwing with 7 veins; Sc and Rs well stalked; M, from upper outer angle of cell; M; and Cu, stalked, from lower outer angle; Cuz from cell very near to M; and Cu.
Male genitalia with uncus in form of pair of triangular plates, each joined on its anterior angle to a large subtriangular process which distally curves medially in tapering to a blunt point. Transtilla present, incomplete. Gnathos with medial process subquadrate in ventral view. Juxta rounded. Vinculum round to subtruncate. Valvae rectangular, apex rounded, inner half somewhat concave. Aedeagus short, compressed, with vesica unarmed.
Female genitalia with apophyses slender, rather straight. Bursa unarmed. Ductus seminalis from posterior end of bursa, slender.
Goya stictella (Hampson), new combination Figures 23, 96, 131 Saluria stictella Hampson, 1918, pp. 96-97. Diacnosts.—This species is easily distinguished from albivenella by the presence of dark brown spots on the forewing and by the absence of well-marked white tracing on the forewing veins.
Description.—Labial palpi porrect, basal segments white, second and third segments light brown on outer sides, third decumbent;
REVISION OF PEORIINAE AND ANERASTIINAE 57
antennae sublaminate in male, light brown, basal segments bearing tuft of scales.
Forewing with M2,; stalked for about one-half their length; ground light brown to grayish red, predominantly white anterior to cell; maculation variable; transverse anterior and median lines marked by dark brown spots in cell; A; with two prominent dark brown spots, one just distal to transverse anterior in the cell, other on the mod- erately well-developed transverse posterior; terminal line of dark brown spots. Hindwing with M; and Cu, long stalked.
Male genitalia with triangular plates of uncus each bearing on its terminal angle a posteromedially directed thorn. Gnathos with medial process bearing short sharp anterior and posterior spines on each lateral margin. Juxta broader than long. Aedeagus tapering strongly posteriorly.
Female genitalia as described for the genus.
Typr.—lIn the British Museum (Natural History).
Type pata.—Lectotype male, hereby designated, labeled as fol- lows: “Type; Bahamas. M.C. Andros. 11. I. 1902 J.L. Bonhote. 1902— 278; Saluria stictella Type o&. Hmpsn.; Pyralidae Brit. Mus. Slide No. 10904.”
SPECIMENS EXAMINED.—5 o’, 2 9.
Distr1IBuTION (Map 4).—In the United States the species is known from Arkansas, Mississippi, and west central Florida.
UNITED STATES: Arxansas: Washington Co., Devel’s Den State Park, 1 o&, May 30, 1966 (R. W. Hodges) [USNM].
Fioripa: Manatee Co., Oneco, 1 o&, 1 9, June 13, 1954 (Paula Dillman) [USNM];1 9, Aug. 3, 1953; Sarasota Co., Siesta Key, 1 &, Apr. 2, 1952 (Charles P. Kimball) [CNC].
Mississippi: Harrison Co., Biloxi, 2 @, June 13, 1917 [CU].
Discussion.—This is the only species of Goya known to occur in the United States; the type locality of albivenella is Argentina.
Subfamily Phycitinae Ragonot
The following seven genera which were previously placed in the Anerastiinae and a new genus, Wakulla, established for Bandera carneella, are transferred to the Phycitinae.
The relationships within the Phycitinae are clear for only a few of the eight genera. Anerastia and Coenochroa show affinities with each other but none with any particular group in the Phycitinae. Barberia is closely related to Anderida, and Bandera to Anagasta. The other four genera have typical phycitine genitalia, although their exact place- ment within the subfamily is uncertain. Of these four, Ragonotia and Martia are closely related and have typical phycitine wing maculation.
5S U.S. NATIONAL MUSEUM BULLETIN 280
Anerastia Hiibner
Anerastia Hiibner [1816]-[1826], p. 367. Ragonot, 1901, pp. 394-397. — Spuler, 1904, p. 200 — McDunnough, 1939, p. 36. [Type: Tinea lotella Hiibner [1810]}-[June 20, 1813.] Designated by Moore, 1886, p. 357.]
Prinanerastia Hampson, 1918, p. 80. [Type: Tinea lotella Hiibner. Original designation. ]
Dracnosis.—Anerastia and two other North American genera possess a toothed frons. In Coenochroa and Martia the teeth are borne at the end of a well-developed protuberance; in Anerastia they form a ring on an otherwise smooth frons.
Description.—Frons rounded, fine projecting serrations forming a transverse elliptical ring about one-half as wide as frons, vesture short; labial palpi deflexed, about 3% times eye length in male, about four times in female; maxillary palpi rudimentary, minute; tongue re- duced; antennae compressed, male shaft with basal segments fused, sublaminate, cilia about one-fifth segment width, female filiform, cilia minute, each segment bearing on its distal half several longer cilia, each about one-half segment width; ocelli well developed.
Forewings with 10 veins; R, from well before upper outer angle of cell; Re free, just before the angle; R; stalked for about one-half its length onto Rs; M, from below the angle; M2,; fused, well separated from Cu,; Cue from well before Cu; at angle. Hindwing with 7 veins; Sc and Rs approximate beyond cell, then diverging; M; and Cu; stalked for over one-half their length, from lower outer angle of cell; Cuz from just below the angle.
Male genitalia with uncus bilobed, membranous medially. Gnathos complex, medial process stout, subrectangular, posteriorly bearing very short truncate lateral arms, mediodorsal process, and strong anteroventrally directed hook, dorsoanteriorly with large curved flat hook between uncus lobes. Transtilla crescent-shaped, poorly developed. Juxta elliptical, anterior margin sclerotized. Vinculum broadly rounded. Valvae tapering on distal third to bluntly pointed apex of costa, anterior-facing crescent-shaped ridge near base of taper- ing portion. Aedeagus with vesica bearing two large cornuti, one lance-shaped, the other deeply bicuspidate.
Female genitalia with ovipositor well sclerotized, tapering to bluntly pointed tip. Dorsum of eighth abdominal segment deeply incised medially. Apophyses well developed, anterior very slightly shorter than posterior. Ductus bursae moderate. Bursa unarmed. Ductus seminalis from posterior end of bursa.
Discusston.—This Old World genus is represented in North America by lotella, a species common in Kurope.
REVISION OF PEORIINAE AND ANERASTIINAE 59
Anerastia lotella Hiibner
Fiaures 34, 48, 64, 97, 132, 167
Tinea lotella Hiibner [1810]-[June 20, 1813].
Anerastia lotella Hiibner [1826], p. 367. — Buckler, 1901, pp. 203-206, pl. 156, fig. 4. — Ragonot, 1901, pp. 397-398. — Spuler, 1904, p. 200, pl. 82, fig. 2. — Beirne, 1952, pp. 74-75, pl. 6, fig. 1.
Prinanerastia lotella (Hiibner). — Hampson, 1918, p. 80.
Anerastia lobella [sic] (Hiibner). — McDunnough, 1939, p. 36.
Draenosis.—This is the only North American species in the genus.
DescripTion.—Labial palpi on outer sides varying brown to light brown, light orange, or orange; antennae orange brown; frons light brown, darker laterally, vertex and occiput brownish white dorsally, occiput laterally, patagia, tegulae, and dorsum of thorax brown to orange brown; legs brown to orange brown on outer sides, brownish white on inner sides.
Forewing varying brown to orange; veins white traced, sprinkled with brown; Az broadly sprinkled with brown; A, fold lightly white traced on outer half. Hindwings brown, fringe light brown.
Genitalia as described for the genus.
Typr.—Lost.
TypE LocaLity.—Not given, but presumably Europe.
RecorpED HostTs.— Poaceae: Ammophila arenaria (L.) Link (European beach grass), larvae feeding on roots near junction with stem; also on Festuca ovina L., Secale cereale L., (Rye) to which they are sometimes a serious pest, and Aira species, (Buckler).
SPECIMENS EXAMINED.— 15207, 459.
DistrisuTion (Map 1).—Alaska (Fort Yukon) south to Washing- ton, east to Minnesota.
UNITED STATES: Atasxa: near Fort Yukon, 1o [USNM].
Minngsora: Cass Co., Cass Lake, 1o June 18, 1934 (A. A. Granovsky) [UM]; 14, June 27, 1934; 247, July 27, 1936 (R. H. Daggy) [UM]; 107, Aug. 2, 1936; Cass Co., 167, June 24, 1937 (L. W. Orr) [UM]; 107, June 26, 1937; 27, June 30, 1937; 14, July 1, 1937 (P. M. Schroeder), 1¢, July 11, 1937 (L. W. Orr); 10, Aug. 11, 1937; Kittson Co., Hallock, 1, Aug. 9, 1935 (D. G. Denning) [UM]; 14, June 20, 1936; 347, June 21, 1936; 267, July 7, 1937; Polk Co., Crookston, 10, July 20, 1935 (D. G. Denning) [UM]; 17, July 21, 1935; 27, July 22, 1935; 16, July 27, 1935; 2, June 21, 1937; 19, June 23, 1937; 307, June 27, 1937; 16, June 26, 1937; 167, June 26, 1941; 1, June 27, 1941.
WasuHineTon: Walla Walla Co., Wallula, 17, July 30, 1948 (William C. Cook) [CNC].
CANADA: ABerta: Banff, 14’, June 29, 1922 (C. B. D. Garrett) [CNC]; Dominion Range Sta. Manyberries, 17, July 10, 1951 (D. F. Hardwick) [CNC]; Scandia, 167, July 9, 1956 (E. E. Sterns) [CNC]; 10’, July 10, 1956.
Mantrosa: Aweme, 1<7, July 26, 1914 (N. Criddle) [CNC]; 307, July 6, 1920; 1c, 19, July 17, 1920; 1c", 49, June 28, 1921; 14, 19, July 1, 1921; 19, (P. N. Vroom);1, 69, July 2, 1921 (N. Criddle) [CNC]; 30, 49, July 6, 1921; 39, July 26, 1921; 1.7, July 28, 1921; 39, Aug. 3, 1921 (P. N. Vroom) [CNC];
285-934—68——__5
60 U.S. NATIONAL MUSEUM BULLETIN 280
16, Aug. 10, 1921 (N. Criddle) [CNC]; 17, Aug. 25, 1921;3¢, 39, June 13, 1922; 29, June 23, 1922; 20, June 27, 1922;19, July 19, 1922;16, July 22, 1922; 20’, Sept. 15, 1923; 10, July 11, 1925; 1¢, Aug. 10, 1925; 19, Aug. 21, 1925; 30, 29, July 2, 1926; 19, Aug. 11, 1926; 7¢, July 20, 1928; Brandon, 1.7’, June 27, 1958 (R. B. Madge) [CNC]; 1¢, July 6, 1958 (R. L. Hurley) [CNC]; 19, July 7, 1958 (R. B. Madge) [CNC]; 1 July 8, 1958;50, 29, July 17, 1958; (R. L. Hurley) [CNC]; 29, July 20, 1957 (R. B. Madge) [CNC]; 1¢7, July 31, 1958 (R. L. Hurley) [CNC]; 19, Aug. 7, 1958 (R. B. Madge) [CNC]; Glenboro, 38%, “Spruce-sand community,” June 17, 1958 (R. L. Hurley) [CNC]; Ninette, 13, “Maple-Elm floodplain community,” June 17, 1958; Wabowden, 1.7, Aug. 7, 1949 (J. B. Wallis) [CNC].
NORTHWEST TERRITORIES: Bathurst Inlet, 19, July 28, 1951 (W. I. Campbell) [CNC]; Hay River, 27, July 30, 1951 (P. R. Ehrlich) [CNC]; 1, July 31, 1951.
Qursec: Forestville, 1¢%, July 11, 1950 (R. deRuette) [CNC].
SASKATCHEWAN: Attons Lake, Cut Knife, 1, June 23, 1940 (A. R. Brooks) [CNC]; 66, July 11, 1940; 27, 19, July 12, 1940; Rutland, 57, July 15, 1940 (A. R. Brooks) [CNC]; 1, Aug. 2, 1940; Saskatoon, 167, June 26, 1923 (Kenneth M. King) [CNC]; 10, July 2, 1923; 24, July 7, 1923; 14, July 21, 1923; Was- kesiu Lake, 19, July 11, 1939 (A. R. Brooks) [CNC]; 27, July 13, 1939; 207, 19, July 15, 1939; 16, July 18, 1939; 2, 19, July 21, 1939.
Coenochroa Ragonot
Coenochroa Ragonot, 1887, p. 20; 1889, p. 117, — Hulst, 1890, p. 217. — Smith, 1891, p. 85. — Ragonot, 1901, pp. 418-419. — Hulst, 1902, p. 441. — Barnes and McDunnough, 1917, p. 150.—Hampson, 1918, p. 58.—McDunnough, 1939, p. 36. [Type: Coenochroa californiella Ragonot, 1887. Original designation. ]
Petaluma Hulst, 1888, p. 116; 1890, pp. 215-216. — Smith, 1891, p. 85. — Hulst, 1902, p. 440. [Type: Anerastia illibella Hulst, 1887. Original designation.]
Alamosa Hampson in Ragonot, 1901, p. 369. — Barnes and McDunnough, 1917, p. 149. — Hampson, 1918, p. 65. — McDunnough, 1939, p. 36. [New synon- ymy.] [Type: Alamosa piperatella Hampson in Ragonot, 1901. Monobasic.]
Diacnosis.—The genus may be easily recognized by the combina- tion of a toothed protuberant frons and longitudinal wing pattern. Martia has a somewhat similar frons, but the wing pattern is predom- inantly transverse. The wing maculation of Anerastia is similar to that of Coenochroa, and the frons is toothed, but not protuberant. The bifid male uncus is unique to Coenochroa.
Derscription.—Frons with prominent anterior protuberance ter- minating in dorsoposteriorly sloping closed rim of irregular teeth sur- rounding central beak (figs. 44-47); labial palpi porrect or deflexed; maxillary palpi rudimentary, naked, usually hidden by labials; tongue reduced, usually hidden by palpi; antennae compressed, each segment bearing pair of perpendicularly projecting cilia ventrally near distal end, male shaft scaled dorsally, ventrally and laterally bearing dense fine recurved cilia, female shaft scaled dorsally and laterally, ventrally with fine sparse recurved cilia; ocelli small, covered with scales.
REVISION OF PEORIINAE AND ANERASTIINAE 61
Forewings with 10 veins; vein Cuz from before lower outer angle of cell; Cu; and M; from angle, stalked; Mz absent; M, from upper outer angle of cell; Ros stalked, from just before the upper outer angle; R,; from well before angle. Hindwing with 6 or 7 veins; vein Cu, from be- fore lower outer angle of cell; Cu: and Ms; from angle, stalked for at least one-half their length, sometimes fused; M2 absent; M, from just below upper outer angle of cell; Rs and Sc from upper angle, stalked for at least one-half their length.
Male genitalia with uncus bifid, setose ventrally, terminating in minute ventrally directed hook. Gnathos with medial process U- shaped. Transtilla absent. Juxta membranous, margin sclerotized anteriorly. Vinculum variable, U-shaped or \V-shaped, anteriorly rounded or bluntly pointed. Valvae broadly rounded, variable in out- line, setose distally. Aedeagus tapering posteriorly, vesica armed.
Female genitalia with ovipositor lobes triangular, moderately setose. Apophyses rather straight and uniform. Ductus bursae moderately broad. Bursa slender and unarmed. Ductus seminalis leaving near mid- point of ductus bursa.
Key to the Species of Coenochroa Based Upon External Structure 1. Central beak of frons protruding beyond rim (figs. 45, 46); costa paler than
cell; R, usually first to branch from stalked Roi,....2.2.2.. illibella Central beak of frons not protruding beyond rim; costa not oe R, usually first to branch from stalked Roi,...... sd
2. Central beak of frons with irregular tip, just pennies eed ee rim ae 47); Atlantic-and Gulf .Coastal Plain 3). s .» «) sy 26a . . . . . bipunctella Central beak of frons pointed and small, not attaining end of rim (fig. 44); Texas and Western United States. ........ ... californiella
Coenochroa californiella Ragonot Ficurses 41, 44, 65, 98, 133, 168
Coenochroa californiella Ragonot, 1887, p. 20; 1889, p. 117. — Hulst, 1890, p.217.— Smith, 1891, p. 85. — Ragonot, 1901, p. 420. — Hulst, 1902, p. 441. — Barnes and McDunnough, 1917, p. 150. — Hampson, 1918, p. 59. — MeDun- nough, 1939, p. 36.
Coenochroa inspergella Ragonot, 1887, p. 20; 1889, p. 117. — Hulst, 1890, p. 216. — Smith, 1891, p. 85. — Ragonot, 1901, p. 419. — Hulst, 1902, p. 440. — Barnes and McDunnough, 1917, p. 150. — Hampson, 1918, p. 59. — McDun- nough, 1939, p. 36. [New synonymy.]
Diaanosis.—This is the only member of the genus in which the central beak of the frons does not attain the rim. The genitalia are identical to those of tlibella, a species which lacks the discal spot and has a distinctive pale costa.
DescripTion.—Frons with protuberance cylindrical, ventrally about as long as wide, rim slanting about 45°, teeth projecting for- ward, central beak a very small cone not reaching rim, vesture light
62 U.S. NATIONAL MUSEUM BULLETIN 280
brown dorsally, white ventrally; labial palpi deflexed, basal segments white, second and third uniform light yellow, occasionally brown dorsally; antennae light brown, finely ciliate; patagia, tegulae, vertex, occiput, and legs varying light yellow to brown.
Forewing radius varying from about 8 mm to 12 mm, average of 9.5 mm; ground color light yellow to orange yellow; veins usually traced with white, often sprinkled with black scales; dark discal spot at lower outer angle of cell; vein Re usually first to leave stalked Rey.
Male genitalia with aedeagus slender, slightly thicker anteriorly, vesica with a single slender conutus.
Female genitalia with ductus seminalis slender, not thickened basally.
Typrs.—C. californiella and C. inspergella, in the Muséum National d’Histoire Naturelle.
Type pata.—C. californiella, lectotype male, hereby designated, Walsingham, genitalia slide No. 536, J. Shaffer, June 20, 1965. In the original description the type locality is given as California.
C. inspergella, lectotype male, hereby designated, Arizona, Morrison, 1881, genitalia slide No. 537, J. Shaffer, June 20, 1965.
SPECIMENS EXAMINED.—260 6, 133 9.
DistriBuTIoN (Map 2).—British Columbia south to California, east to Kansas and Texas.
UNITED STATES: Arizona: Cochise Co., Carr Canyon, Huachuca Mts., 2 o', June 3, 1952 (M. Cazier, W. Gertsch, R. Schrammel) [AMNH]; Ft. Grant, Pinaleno Mts., 1 9, July 15-19, 1917 (C. U. Biol. Expend.) [CU]; Huachuca Mts., 1 o&, May 1-7 [USNM]; 1 2, August 8-15; Montezume Pass, Huachuca Mts. (6500 ft.), 1 o&, Sept. 7, 1950 (T. Cohn, P. Boone, M. Cazier) [AMNH]; Paradise, 1 o’, March [USNM]; Paradise, 1 9, July (Poling) [CM]; 1 @&, 3 9, August; Portal, 1 9, June 1, 1952 (M. Cazier, W. Gertsch, R. Schrammel) [AMNH]; San Bernardino Ranch (3750 ft.), 1 9, August (F. H. Snow) [UK]; Southwestern Research Sta., 5 mi. west of Portal (5400 ft.), 1 o&, Apr. 10, 1956 (Cazier, Ordway) [AMNH]; 1 o, Apr. 22, 1956; 1 o&, Apr. 23, 1956; 1 &, Apr. 25, 1956; 2 co, May 16, 1956 (M. Statham); 1 o&, 2 ?, May 18, 1956; same locality, 1 9, July 28, 1959 (HE. G. Linsley) [UCB]; same locality, 1 9, July 26—Aug. 3, 1959 (A. B. Klots) [ABK]; same locality, 1 9, Nov. 1, 1959 (J. R. Powers) [UCB]; 1 o&, Nov. 2, 1959; same locality, 1 #@, May 16, 1960 (Carl W. Kirkwood) [CPK]; 1 o, May 18, 1960; 1 9, May 25, 1960; 1 o&, May 18, 1960; 1 9, May 25 1960; 1 &, May 28, 1960; 1 o&, May 29, 1960; 1 o&, Apr. 10, 1961; 1 &, Apr. 12, 1961; 1 o&, Apr. 13, 1961; 1 o, Apr. 17, 1961; same locality, 1 9, Apr. 23, 1961 (M. A. Cazier) [AMNH]; 1 o&, Apr. 25, 1961; 1 9, May 2, 1961; same locality, 1 o, 2 2, May 10, 1961 (Gertsch & Cazier) [AMNH];1 &, 1 9, May 15, 1961 (M. Cazier); 1 @, May 27, 1961 (M. Statham); same locality, 1 &, Apr. 10, 1962 (Carl W. Kirkwood) [CNC]; 4 o&, Apr. 15, 1962; 1 &, Apr. 16, 1962; 1 o', Apr. 18, 1962; 1 o&, Apr. 19, 1962; 2 &, Apr. 20, 1962; 1 &, Apr. 21, 1962; 2 o', Apr. 22, 1962; 1 o&, Apr. 23, 1962; 1 o&, Apr. 27, 1962; same locality, 1 o, Aug. 23, 1962 (M. Statham) [AMNH]; 1 o&, Aug. 30, 1962; Coconino Co., Flag- staff, 2 &, no date ( H. 8. Barber) [USNM]; Gila Co., vicinity Globe, 3 9, Aug. 4-5, 1937 (A. B. Klots) [AMNH]; Globe, 1 o&, May 18, 1950 (E. C. Johnston)
REVISION OF PEORIINAE AND ANERASTIINAE 63
[CNC]; San Carlos, 1 o, May 12-13 (1918 (J. C. Bradley) [CU]; no locality, 1 9, no date (O. C. Poling) [USNM]; Maricopa Co., New River, 1 #, May 7, 1950 (E. C. Johnston) [CNC]; Tempe, 1 o’, Apr. 19, 1920 (E. V. Walter and H. L. Arnold) [USNM]; 1 o, Apr. 26, 1920; 1 9, May 3, 1920; 1 9, July 26, 1920; 1 9, Sept. 27, 1920 (E. V. Walter); Pima Co., Baboquivari Mts. (5-7000 ft.), 1 &, July 15-30, 1923 (O. C. Poling) [USNM]; 1 9, July 27-31, 1923; 1 o', 1 9, Aug. 1-4, 1923; 3 9, Aug. 1-15, 1923; 3 o, 7 9, Aug. 15-30, 1923; 1 9, Sept. 1-15, 1924; 1 @, Sept. 15-30, 1924; 11 #, 12 9, Oct. 1-15, 1924; 9 o, 1 9, Oct. 15-30, 1924; 1 o&, Nov. 1-15, 1924; Baboquivari Mts., 3 3, 2 9, Apr. 23, 19388 (J. A. Comstock) [LACM]; 4 o&,5 9, Apr. 24, 1928; 14 ¢, 1 9, Apr. 25, 19388; 1 o&, 2 9, Apr. 26, 1938; Baboquivari Mts., 1 9, no date (F. H. Snow) [UK]; Madera Canyon, Santa Rita Mts. (4400 ft.), 2 3, 2 9, May 26, 1963 (J. G. Franclemont) [JGF]; 1 9, June 2, 1963; 1 9, June 4, 1963; Pinal Co., Oracle, 1 9, May 19, 1933 (Grace H. and John L. Sperry) [AMNH];1 2, June 4, 1935, [USNM]; [CUC];1 9, [AMNH]; 3 9, June 5, 1935 [AMNH}]; Boyce Thompson Arboretum, Superior, 1 9, Aug. 1, 1937 (A. B. Klots) [AMNH]; Santa Curz Co., Madera Canyon, Santa Rita Mts., 1 o, Aug. 15, 1949 (Lloyd M. Martin) [LCAM]; (5800 ft.), 1 9, June 22, 1955; Madera Canyon, Santa Rita Mts., 1 o@, Aug. 24-Sept. 2, 1957 (William A. Hammer) [LACM]; same locality (4880 ft.), 1 o&, Apr. 14, 1963 (J. G. Franclemont) [JGF]; 1 0, May 20, 1963; 1 o', May 21, 1963; 1 9, May 28, 1963; 1 9, June 1, 1963; Santa Rita Mts., 4 9, Aug. 22, 1946 (J. A. Comstock and Lloyd M. Martin) [LACM]; 2 9, Aug. 24, 1946; 1 9, Aug. 27, 1946; 1 9, Aug. 29, 1946; Nogales, 1 9, May 30, 1899 [USN M]; Nogales, 2 o&, 3 9, May 24-30 [USNM];1 0,2 9, June 1-7; 1 o&, July 8-15; Pena Blanca (3950 ft.), 1 9, June 7, 1963 (J. G. Franclemont) [JGF]; Yavapai Co., 10 mi. east of Congress, 1 9, Aug. 22-23, 1927; Mayer, 3 o&, 1 9, May 21, 1959 (M. O. Glenn) [MOG]; Prescott, 6 o, May 6, 1950 (E. C. Johnston) [CNC]; no county, ‘‘Catal Spgs,” 2 9, (E. A. Schwarz) [USNM]; Fish Creek, Tonto Nat. For., 5 @, 3 9, May 9-10, 1918 (J. G. Bradley) [CU]; Todd’s Lodge, Oak Creek Canyon, 1 <o, June 12, 1941 (Grace H. and John L. Sperry) [CNC]; Santa Rita Mts., 1 #7, 1 9, May 21, 1898 (E. A. Schwarz) [USNM]; 1 9, May 26, 1898; 1 o, June 8, 1898; 1 9, June 13 1898; 1 9, June 14, 1898; Santa Rita Mts. (5-8000 ft.), 1 @, June (F. H. Snow) [UK]; no locality, 1 @, no date (Morrison), labeled “Coll. Ragonot 95-85” [BM]; no locality, 1 o&, 1881 (Morrison) [USN M]; no locality, 1 @, August (O. Poling) [USNM]; 6 <&, September (Poling) [CM]; 1 o, no date, from Fernald collection [USNM]; 1 <, no date, from C. V. Riley collection [USNM];1 0,1 9, no date [USNM].
CairornNia: Inyo Co., Bishop, 2 o, June 14, 1937 (E. C. Johnston) [CNC]; Lassen Co., Ravendale, 1 9, June 20, 1959 (G. I. Stage) [ABK]; Modoc Co., Canby, 3 ao’, July 16, 1936 (E. C. Johnston) [CNC]; Orange Co., Fullerton, 2 9, Nov. 14, 1962 (C. A. Toschi) [UCB]; Riverside Co., Rancho La Sierra, Arlington, 1 co, Aug. 26, 1949 (A. H. Rindge) [AMNH]; 1 o&, Aug. 23, 1952; Idyllwild, 1 @, May 13, 1937 (H. Little) [AMNH]; Lake Hemet, 6 &, June 9, 1937 (E. C. Johnston) [CNC]; Riverside, 1 @, May 4, 1926 (Grace H. and John L. Sperry) [CPK]; 1 7, Sept. 28, 1931 (C. H. Dammers) [USNM]; 1 &, 1931; 1 9, May 4, 1934 (Grace H. and John L. Sperry) [CPK]; 2 &@, May 23, 1935 (Grace H. and John L. Sperry) [AMNH], [CNC]; 1 o&, 1 9? [AMNH]; 1 @, July 12, 1985 (Grace H. and John L. Sperry) [AMNH]; 1 o, Apr. 29, 1936 [CNC]; 1 o, Apr. 27, 1937 (H. Buckwalter) [AMNH]; 3 2, June 1, 1939 [AMNH]; San Bernardino Co., Barton Flats, 2 @, June 27, 1946 [AMNH]; Upper Santa Ana River, 1 9, June 2, 1946 [AMNH]; 1 o, July 10, 1946; 1 &, July 18, 1946;
64 U.S. NATIONAL MUSEUM BULLETIN 280
1 o, Aug. 18, 1948 (Melander); San Diego Co., Mt. Palomar, 3 o’, July 18, 1963 (J. Powell) [UCB]; Mt. Palomar St. Pk., 1 o&, July 12, 1953 (W. J. and J. W. Gertsch) [AMNH]; San Diego, 1 o, June 16, 1912 (George H. Field) [ASNM]; Julian, 2 ?, no date [USNM]; Siskiyou Co., Indian Butte, 1 @, July 16, 1936 (KE. C. Johnston) [CNC]; county unknown, Hathaway Creek, San Bernardino Mts., 2 o&, Aug. 2, 1940 (J. A. Comstock and C. Henne) [LACM].
Ipano: Canyon Co., Parma, 1 o, July 6, 1951 (A. J. Walz) [AMNH].
Nevaba: Clark Co., Kyle Canyon, Charleston Mts., 1 9, Apr. 26, 1950 (E. C. Johnston) [CNC]; Wheeler’s Spring, Charleston Mts., 2 o&, May 14, 1934 [CPK].
New Mexico: Bernalillo Co., Albuquerque, 1 ¢ , July 21, 1902 (Oslar) [USNM]; 1 9, July 24, 1902; 1 &, 9, no date [UK]; Hidalgo Co., Rodeo, 1 9, Nov. 7, 1959 (J. R. Powers) [UCB]; county unknown, ‘‘so. N. Mex.,” 1,1 9, Aug. 23-30 (Poling), Rothschild Bequest B.M. 1939-1 [BM].
Texas: Blanco Co., 1 9, no date, G. D. Hulst [AMNH]; Brewster Co., Alpine, 1 9, May 22, 1950 (EK. C. Johnston) [CNC]; Burnet Co., 2 o&, 2 9, no date (F. G. Schaupp) [USNM]; Carmeron Co., Brownsville, 2 9, May 31, 1904 (H. 8S. Barber) [USNM]; 1 9, June 8, 1904; 1 o&, June [UK]; 2 o&, June [ANS]; Hidalgo Co., Mercedes, 1 9, Aug. 31, 1958 (H. Smalzried) [AMNH]; 1 o, no date [CPK]; Jeff Davis Co., Limpia Canyon, 8 o&, 2 ?, May 20, 1950 (E. C. Johnston) [CNC]; Randall Co., Palo Duro Canyon St. Pk., 3 &, May 5, 1961 (Lloyd M. Martin, Robert H. Reid, William A. Rees, Robert J. Ford) [LACM]; 3 o', May 6, 1961; 3 o, May 8, 1961; 1 9, May 9, 1961; 3 o, 1 9, May 11, 1961; 1 9, May 12, 1961.
Urau: Utah Co., Vineyard, 2 o’, July 7, 1917 (Tom Spalding) [ANS]; 1 2, July A) 197-1 ot, Aug.i6; 1917.
Wasuineton: Adams Co., Othello, 1 o&, June 23, 1959 (R. F. Harwood) [USNM]; 1 o&, July 9, 1959; Chelan Co., First Creek, 2 @, July 2, 1949 (E. C. Johnston) [CNC]; Grant Co., Dry Falls, 1 &@, June 30, 1949 (E. C. Johnston) [CNC]; Quincy, 1 &, July 5, 1959 (C.S. Crawford) [USNM]; Okanogan Co., Black Canyon, 4 o&, July 1, 1949 (E. C. Johnston) [CNC]; Walla Walla Co., Walla Walla, 1 o&, June 27, 1935 (H. P. Lanchester) [USNM]; Whiteman Co., Snake River, opposite Clarkston, 1 @, May 29, 1931 (J. F. Clarke) [USNM]; Yakima Co., Yakima, 1 <&, June 8, 1931 (Fred Dauy) [USNM]; 1 2, Sept. 11, 1955 (A. I. Good) [AMNH].
CANADA: British CoLumsia: Kamloops, 1 9, June 28, 1937 (J. K. Jacob) [CNC]; Oliver (1500 ft.), 1 o&, June 5, 1953 (D. F. Hardwick) [CNC]; 7 o, June 16, 1953; 13 o&, June 17, 1953; (2500 ft.), 3 #, June 26, 1953; (1000 ft.), 1 o&, July 9, 1953 (J. E. H. Martin) [CNC]; (2500 ft.), 2 @, July 10, 1953 (D. F. Hardwick) [CNC]; Osoyoos, 1 &, May 19, 1923 (C. B. Garrett) [CNC]; (1200 ft.), 1 9, July 22, 1953 (D. F. Hardwick) [CNC].
Discusston.—The degree of black scaling on the forewings is quite variable and ranges from a very marked broad tracing of the veins to an almost total absence of black scales. The type specimen of californiella is very sparsely set with black scales and that of insper- gella rather heavily so, thus it is not surprising that they were regarded as representing separate species. Wing venation, frons, and genitalia are essentially identical in the two holotypes.
Small specimens may easily be mistaken for bipunctella.
REVISION OF PEORIINAE AND ANERASTIINAE 65
Coenochroa illibella (Hulst)
Fiaures 39, 40, 45, 67, 134, 169
Anerastia illibella Hulst, 1887, p. 188.—Rindge, 1955, p. 164.
Coenochroa puricostella Ragonot, 1887, p. 20.
Coenochroa illibella (Hulst).—Ragonot, 1889, p. 117; 1901, p. 419.—Barnes and McDunnough, 1917, p. 150.—Hampson, 1918, p. 58.—McDunnough, 1939, p. 36.
Alamosa piperatella Hampson in Ragonot, 1901, pp. 369-370.—Barnes and McDunnough, 1917, p. 149.—Hampson, 1918, p. 65.—McDunnough, 1939, p. 36. [New synonomy.]
Diacenosis.—The species may be recognized by either the pale costal region of the forewing, the compressed and protruding central beak of the frons, the absence of a discal spot, or the fact that Re is almost always the first vein to leave the stalked Ro,..
Description.— rons with protuberance cylindrical, ventrally about as long as wide, rim slanting at about 45°, teeth projecting forward, central beak strongly compressed, protruding well beyond rim, vesture light brown dorsally, white ventrally; labial palpi some- what deflexed, basal segments white, second and third brown on outer sides and dorsally, light brown to white ventrally; antennae as in californiella; patagia, tegulae, vertex, occiput, and legs as in cali- forniella.
Forewing radius averaging about 8.8 mm; area anterior to cell white, sometimes variously sprinkled with black scales; region posterior to radius quite variable, usually grayish orange sprinkled with black scales predominantly on veins; area between veins sometimes white, veins never traced with white; vein R: first to branch from Rois. Hindwing with fringe white.
Genitalia as in californiella.
Typrs.—A. illibella, in the American Museum of Natural History; C. puricostella, in the Museum National d’Histoire Naturelle; A. prperatella, in the British Museum (Natural History).
Type pata.—A. illibella, lectotype female, hereby designated, Blanco Co., central Texas, November, collection G. D. Hulst, geni- talia slide No. 3227, Carl Heinrich, June 14, 1946. In the original description Hulst reports only that he had two males and one female from Texas.
C. puricostella, lectotype male, hereby designated, Arizona, Morison, genitalia slide No. 538, J. Shaffer, June 20, 1965.
A. piperatella, lectotype male, hereby designated, labeled as follows: “Syntype; Colorado Ameriq. sept.; Coll. Ragonot, 95—85.; Alamosa piperatella Rag. Colorado ex Coll. Rag; o@ genitalia slide I-4-1967 J. Shaffer No. 706.”
SPECIMENS EXAMINED.—104 o’, 569.
66 U.S. NATIONAL MUSEUM BULLETIN 280
DistrriBuTION (Map 2).—Ontario west to Alberta and Washington, south to California and Texas.
UNITED STATES: Arizona: County unknown, “Br’t Angel,’ 1 <, no date [USNM].
Cauirornia: Inyo Co., 7 mi. north Parcher’s Camp, 4 o, June 30, 1961 (J. Powell) [UCB]; Mono Co., Leevining, 2 @, July 19, 1988 (E. C. Johnston) [CNC]; 1 &, July 20, 1938; 1 mi. southwest Tom’s Place, 1 7,2 9, Aug. 10, 1963 (C. A. Toschi) [UCB]; 1 @, Aug. 13, 1963 (M. J. Tauber and C. A. Toschi).
Cotorapo: Adams Co., Watkins, 8 o, July 4-5, 1927 [CU]; Boulder Co., Boulder, 1 &, June-August, 1896 [BMNH]; Chaffee Co., Salida, 1 #, July 9, 1937 [AMNH]; Denver Co., Denver, 1 &, no date [USNM]; 2 &, (Oslar); Prowers Co., Lamar, 2 o’, Sept. 24, 1945 (E. C. Johnston) [CNC]; county unknown, 1 o, no date (Oslar) [USNM]; 1 o, no date [AMNH]; 6 o@, no date [USNM].
Kansas: Clark Co. (1962 ft.), 1 9, June (F. H. Snow) [ANS]; 1 @ [UK]; 2 9 [USNM].
New Mexico: Eddy Co., Artesia, 2 &, May 13, 1950 (E. C. Johnston) [CNC] Sandoyal Co., Frijoles Canyon, Bandelier Nat. Mon., 1 &, July 9-11, 1957; (A. B. Klots) [AMNH]; (6050 ft.), 4 @#, July 17, 1962 (E. and I. Munroe) [CNC]; Otero Co., Mescalero, 1 o, May 12, 1950 (E. C. Johnston) [CNC]; Dona Ana Co., Mesilla, 1 9 , July 30 (Cockerell) [BM]; Mesilla Park, 1 7, 1 9, May 8 (Cockerell) [USNM]; 3 o&, 3 9, May 13; 2 o, (3800 ft.) [CNC]; 1 #, May 22 (Cockerell) [CNC]; 1 o« [USNM]; 1 9, June 8; 1 9, July 8;1 9, Aug. 29; 1 9, no date; 1 9? [CNC]; Mesquite near Mesille Park, 1 9, July 12, 1917 [CNC]; 5 o& [CU]; Grant Co., Silver City, 1 o&, May 25, 1913 (J. B. Wallis) [CNC]; 1 9, May 27 1913; 1 9, July 25, 1913; 4 6, 3 9, July 26, 1913; 2 o, July 27, 1913; 7 o#, 1 9, July 28, 1913; county unknown, southern New Mexico, 2 ?, Aug. 23-30 (Poling) [BM].
Norto Daxora: Ransom Co., 1 mi. southeast McLeod, 1 @, July 19, 1963 (J. R. Powers) [UCB].
SoutH Daxora: Davison Co., Mitchell, 1 @, Sept. 14, 1945 (E. C. Johnston) [CNC].
Trexas: Reeves Co., Pecos, 2 o&', 17 9, May 18, 1950 (E. C. Johnston) [CNC]; 1 &, 8 9, June 2, 1950; county unknown, 1 9, no date, labeled, ‘‘Fernald Col- lection,” “U.S.N.M. Type No. 40077,” ‘Petaluma illibella Hulst, Type,’’ “Anerastia illibella Hulst, Type’? [USNM].
Urau: Garfield Co., 11 mi. southeast Panguitch (7200 ft.), 4 &#, July 11, 1960 (F., P., and B. Rindge) [AMNH]; 3 2, July 12, 1960; 5 &, July 13, 1960; Tooele Co., Stockton, 1 9, Aug. 8, 1904 (Tom Spalding) [USNM]; 1 9, no date; Utah Co., Vineyard, 1 o, Aug. 6, 1917 (Tom Spalding) [ANS]; 1 9, July 16, 1917.
WASHINGTON: Grant Co., Quincy, 1 &, July 1, 1959 (C. S. Crawford) [USNM]; 1 &, July 5, 1959.
CANADA: Avserta: Medicine Hat, 1 0, July 15, 1956 (E. E. Sterns) [CNC]; Scandia, 1 6, July 6, 1956 (EK. E. Sterns) [CNC].
Onrario: Marmora, 1 9, Aug. 16, 1952 (J. F. McAlpine) [CNC]; Point Pelee 1 o,1 9, June 29, 1927 (F. P. Ide) [CNC].
Discussion.—The original description of illibella lists two males and one female from Texas. The disposition of the male types is not known, and two female specimens from Texas bear Hulst’s ‘“Type” label. Available evidence is insufficient to determine which, if either, of these two females was included in the original series, and I have
REVISION OF PEORIINAE AND ANERASTIINAE 67
designated as lectotype the specimen in the American Museum of Natural History. The remaining female is deposited in the U.S. National Museum, full data being given above.
The specimens from Pecos, Tex. are atypical in that the frons beak is unusually well developed and the rim is flared anteriorly so that the teeth point somewhat anterolaterally. Most of the Pecos speci- mens are only sparsely set with black scales on the forewings, but otherwise are typical of the species.
Coenochroa bipunctella (Barnes and McDunnough), new combination Figures 42, 47, 66, 135, 170 Alamosa bipunctella Barnes and McDunnough, 1913, p. 184; 1917, p. 149. — Grossbeck, 1917, p. 134. — Hampson, 1918, p. 65. — McDunnough, 1939, p. 36. — Kimball, 1965, p. 251.
Draenosis.—This is the only member of the genus having numer- ous small cornuti on the vesica of the male rather than a single one. The ductus seminalis of females is thickened where it leaves the ductus bursa, not slender as in other species. Otherwise, specimens could be confused with small examples of californiella, which has similar wing venation and maculation, but the two species are largely or wholly allopatric.
DescripTion.—Frons with protuberance cylindrical, ventrally about as long as wide, rim slanting at about 45°, teeth projecting forward, central beak conical with irregular tip even with end of rim, vesture light brown to brown dorsally, white to light brown ventrally; labial palpi usually porrect, basal segments white, second and third brown dorsally and on outer sides, light brown ventrally; antennae as in californiella; patagia and tegulae brown, vertex and occiput light brown dorsally, occiput brown dorsolaterally, white below middle of eye; legs brown on outer sides, white on inner sides.
Forewing radius averaging about 6.9 mm; cell, discocellular, radials, medials, cubitals, and anal veins each traced with a broad white band sprinkled with black scales; area anterior to cell varying yellow to brown and sprinkled with black scales; ground color yellow to yellow brown; prominent black discal spot at lower outer angle of cell; brown on underside; vein R, first to branch from stalked R24. Hindwing bordered with double brown line; fringe white; ground color light brown, darker in apical area; Cu; and M; stalked for at least one-half length, sometimes fused.
Male genitalia with vinculum V-shaped, pointed anteriorly. Aedeagus club-shaped vesica bearing numerous small cornuti which diminish in size anteriorly.
Female genitalia with ductus bursae rather short. Ductus seminalis thickened basally.
68 U.S. NATIONAL MUSEUM BULLETIN 280
Typr.—lIn the U.S. National Museum.
Typr pata.—One female, Fort Myers, Fla., Apr. 16-23, Barnes collection; USNM 69383; genitalia slide No. 573, J. Shaffer, Nov. 23, 1965.
SPECIMENS EXAMINED.—63 o', 39.
DistrisuTion (Map 2).—Atlantic and Gulf Coastal Plain, recorded from Mississippi to New Jersey.
UNITED STATES: Fioripa: Alachua Co., Gainesville, 1 9, June 2, 1927 (J. Speed Rogers) [CU]; 1 o&, June 29, 1927; 1 o&, July 7, 1927;8 o&, July 8, 1927; 1 of, 19, July 10, 1927;1 # [CNC]; 5 o&, July 1927 (J. Speed Rogers) [CU] 1 0, Apr. 20, 1952 (O. Peck) [CNC]; 1 &, Apr. 22, 1952; Dade Co., South Campus, Richmond, 2 o, Nov. 27, 1951 (F. G. Butcher) [CNC]; Indian River Co., Vero Beach, 1 o, no date (J. R. Malloch) [USNM]; Orange Co., Orlando, 1 o, June 14-18, 1927 (C. C. McBride) [CU]; Pasco Co., Elfers, 1 9, Apr. 17, 1952 (G. 8. Walley) [CNC]; 6 o&, Apr. 18, 1952 (J. R. Vockeroth) [CNC]; Polk Co., Lake Alfred, 1 o&, June 30, 1928 (lL. J. Bottimer) [USNM]; Sarasota Co., 1 o&', May 5, 1946, (C. P. Kimball) [CPK]; Siesta Key, 1 o, Feb. 23, 1951 (C. P. Kimball) [CNC]; 10, Feb. 27, 1951; 1 o&, Dec. 24, 1941; 2 o, Feb. 13, 1952; 1 o&, Feb. 28, 1952;3 o, Mar. 1, 1952;1 o&, Mar. 26, 1952;2 o, Apr. 2, 1952; 1 o&, Nov. 6, 1952; 1 o', Nov. 15, 1952; 1 &, Apr. 11, 1953; 1 o, Apr. 24, 1953; 1 o, Feb. 13, 1954; 1 o', Mar. 14, 1954; 1 of, Feb. 9, 1955; 1 o, Feb. 18, 1955; 1 o&, Apr. 29, 1956; 1 6, Oct. 24, 1956; 1 9, Jan. 23, 1957; Seminole Co., Altamont, 1 o’, Sept. 21, 1924 (F. R. Cole) [USNM].
MississipP1: Forrest Co., Camp Shelby, 1 o, Sept. 1-15, 1944 (C. D. Michener) [AMNH].
New JERSEY: Ocean Co., Lakehurst, 1 o&, July 24, 1939 (E. P. Darlington) [ANS].
Norta Carouina: Robeson Co., Maxton, 1 o', May 11, 1944 (A. B. Klots) [AMNH]; 2 o&, May 23, 1944; 1 o, Oct. 4, 1944.
Discusston.—The species has been poorly collected outside of Florida, as is generally true for Coastal Plain groups, and is probably more common there than the few scattered records indicate. It may be sympatric with californiella in the Coastal Plain section of Texas.
Bandera Ragonot
FIGURE 68
Bandera Ragonot, 1887, p. 19. — Hulst, 1890, p. 202 [listed in Phycitinae].— Smith, 1891, p. 84 [listed in Phycitinae].— Ragonot, 1889, p. 117; 1901, p. 409. — Hulst, 1902, p. 440. — Barnes and McDunnough, 1917, p. 150. — Hampson 1918, p. 89. — McDunnough, 1939, p. 36. — Heinrich, 1956, pp. 1, 315, 316. [Type: Anerastia binotella Zeller, 1872. Original designation.]
Nasutes Hampson, 1930, p. 53 [described in Phycitinae]. — McDunnough, 1939, p. 28. — Heinrich, 1956, pp. 315-316 [listed as synonym of Bandera]. [Type: Nasutes venata Hampson, 1930, Monobasic.]
Diacnosis.—The absence of ocelli and the presence of a tongue, reduced but exposed between the palpi, serve to distinguish this genus from the other North American Phycitinae.
REVISION OF PEORIINAE AND ANERASTIINAE 69
Description.—Frons conical; labial palpi porrect, outer sides of second and third segments clothed with white-tipped brown scales, basal segments clothed with broad white scales; maxillary palpi cylindrical, almost reaching frons; tongue exposed between palpi; antennae filiform, compressed, ciliate ventrally; ocelli absent; legs brown to fuscous laterally, tarsi brown and speckled with white.
Forewings with a brown-speckled white band anterior to cell; veins traced with white and margined on both sides with a sprinkling of brown scales; 9 veins; R, from well before outer angle of cell; R, from cell; Ri,s5 fused; M, from upper outer angle; M,,; fused and from the angle; Cu, from before the angle; Cu, from well before the lower outer angle of cell. Hindwing light brown above and below, slightly darker toward apex; 7 veins; Sc and Rs long stalked; Cu, stalked with fused M.,3; Cuz from well before the lower outer angle of cell.
Male genitalia with uncus broadly triangulate, slightly tapering, with apex broadly rounded, laterally and dorsocaudally setose. Gnathos arms terminating in an anterior ventral hook; apical process a simple posteriorly directed hook. Juxta U-shaped, with small lateral knobs. Vinculum stout. Valvae distally rounded; costa tubular, slightly tapering distally. Vesica unarmed.
Female genitalia with ovipositor tapering distally, about as long as basal width. Apophyses slender, well sclerotized; posterior about five or six times as long as their basal separation; anterior about two to four times as long as their basal separation. Ductus bursae long, weakly sclerotized. Bursa small; with a group of three or four small barlike signa near posterior end. Ductus seminalis threadlike; from near or at posterior end of bursa near signa.
Discussion.—The genus shows affinities to a number of other phycitine genera and is probably closest to Anagasta Heinrich. The ereat preponderance of males in collections suggests that females are far less readily attracted to light.
Key to the Species of Bandera Based Upon the Maculation
1. Space between vein A, and cell of forewing a pale yellow field sprinkled with
brown or with brown scales on fold of Ay .... . . . . . Virginella Space between vein A, and cell of forewing a solid y oMewy field, not sprinkled with brown. .. . Be gt ae Ct) egy oe Aa ae Semel Snes
2. Basal area of costa vollaw sas . . . cupidinella Entire length of costa white, eormeted mie ocean genes . . . binotella
Bandera binotella (Zeller) Ficures 31, 68, 99, 136, 171
Anerastia binotella Zeller, 1872, pp. 554-555. Bandera subluteella Ragonot, 1887, p. 19; 1889, p. 117. — Smith, 1891, p. 84 [listed in Phycitinae]. — Ragonot, 1901, pp. 410-411. — Hulst, 1902, p.
70 U.S. NATIONAL MUSEUM BULLETIN 280
440. — Barnes and McDunnough, 1917, p. 150. — Hampson, 1918, p. 90. — McDunnough, 1939, p. 36. [New synonymy.]
Bandera binotella (Zeller). — Ragonot, 1889, p. 117. — Hulst, 1890, p. 202. — Smith, 1891, p. 84. — Ragonot, 1901, pp. 409-410. — Hulst, 1902, p. 440. — Dyar, 1908¢, p. 116. — Barnes and MeDunnough, 1917, p. 150. — Hampson, 1918, pp. 89-90. — McDunnough, 1939, p. 36.
Draenosis.—The presence of a transverse posterior line of dots and a transverse anterior dot on vein A, of the forewing, a truncate vinculum, and an incomplete transtilla each by itself distinguish this species from other members of the genus.
Description.—Frons about as long as diameter of eye, light brown; vertex, occiput, tegula, and prothorax white to light brown dorsally, gray brown laterally; abdomen light creamy brown dorsally, brown ventrally.
Forewing yellow between veins; discal cell, area between cell and vein A,, and area posterior to A, solid yellow; transverse posterior variously developed as a row of dots on veins, best represented by a dark spot near end of vein A,; a dark spot on vein A, halfway between tp line and wing base; light brown to brown on underside.
Male genitalia with transtilla incomplete. Lateral knobs of juxta well developed, setose. Vinculum rectangular, broadly truncate ante- riorly. Aedeagus tapering posteriorly, subtruncate anteriorly. Eighth abdominal segment of male bearing lateral hair tufts.
Female genitalia with ovipositor sparsely setose. Ductus bursae broad.
Typres.—A. binotella, in the British Museum (Natural History), (from Zeller collection); B. subluteella, in the Muséum National d’Histoire Naturelle.
Type pata.—A. binotella, in the original description Zeller reports ‘“Vaterland: Texas. Belfrage fing das Exemplar am 12. Juli.”” Type examined by Mr. Paul Whalley.
B. subluteella, lectotype male, hereby designated, labeled as follows: “TYPE; WALSM; 1901 coll. E. L. Ragonot Museum Paris; Bandera subluteella Rag. type orig. pl. XL f. 21; @ genitalia slide July-28- 1965 J. Shaffer No. 553.’”’ The specimen also bears a small handwritten label inscribed with “‘Co19,” probably intended to be “Colo” (Colo- rado). Ragonot may have interpreted the inscription as an abbrevia- tion for California since that state is given as the type locality. I have seen no specimens of binotella from California although the species is found in Washington and therefore might be expected to occur in northern California.
SPECIMENS EXAMINED.—71 7,8 Q.
DistrinuTIon (Map 4).—Washington southeastward to Arizona and ‘Texas; eastern population on Atlantic Coastal Plain, New Jersey to Massachusetts.
REVISION OF PEORIINAE AND ANERASTIINAE tl
UNITED STATES: Arizona: Apache Co., Greer, White Mts. (8500 ft.) 1 o, Aug. 6, 1962 (E. and I. Munroe) [CNC].
Cotorapo: Rocky Mountain National Park, 9 @, Aug. 15, 1937 (A. B. Klots) [AMNH].
Connecticut: New Haven Co., East River, 1 o&, July 10, 1909 (Charles R. Ely) [USNM]; 1 o, July 30, 1910; 1 @, Aug. 13, 1910; 1 @, Aug. 29, 1910; 1 o&, August 1910; 1 9, July 1911; Windham Co., Putnam, 1 0, Aug. 9, 1940 (A. B. Klots) [AMNH]; 1 9, August 1942; 1 #, Aug. 16, 1960 [ABK].
MassacuvseEtTts: Barnstable Co., Barnstable, 1 9, July 4, 1949 (C. P. Kim- ball) [CU]; same locality, 1 &, Sept. 11, 1949 (C. P. Kimball) [CPK]; 2 2, Aug. 5, 1950; 1 &, Sept. 12, 1952; Dukes Co., Martha’s Vineyard, 1 9, July 20 (F. M. Jones) [CPK]; Worcester Co., Worcester, 1 o, July 11, 1933 [CU].
Montana: Roosevelt Co., Poplar, 1 @, July 15, 1921 (H. G. Dyar) [USNM].
New Jersey: Bergen Co., Oakland, 1 ? , July 26, 1947 (C. P. Kimball) [CNC]; 3 o', Aug. 4, 1948; 3 o, Aug. 6, 1948; 1 9, Aug. 8, 1948 [CU]; 1 o&, Aug. 8, 1948 [CNC]; 3 o, Aug. 9, 1948 [CNC, CPK]; 2 o&, Aug. 10, 1948 [CPK]; 3 &, Aug. 13, 1948 [CNC, CPK]; 2 o, Aug. 14, 1948 [CPK]; 1 @, Aug. 22, 1948 [CPK];1 7, 1 9 , Aug. 24, 1948 [CNC]; 1 o, August 1948; “SM Beach,” 1 o, “7.2”? (F. Haim- bach) [USNM].
New Mexico: Colfax Co., Cimarron Canyon, Sangre de Cristo Mts. (7900 ft.), 1 o, July 7, 1962 (E. and I. Munroe) [CNC]; 1 ¢@, July 13, 1962; Lincoln Co., Cedar Creek Camp, 2 mi. north of Ruidoso (7000 ft.), 1 @, June 30, 1961 (F., P., and J. Rindge) [AMNH]; 6 a, July 29, 1962 (KE. and I. Munroe) [CNC]; Mc- Kinley Co., McGaffey, Zuni Mts. (7500 ft.), 2 @, July 20, 1962 (E. and I. Munroe) [CNC]; 4 o, July 21, 1962; 2 &, July 22, 1962; 2 o&, July 24, 1962; Sandoyal Co., Frijoles Canyon, Bandelier Nat. Mon. (6000 ft.), 1 o&, July 17, 1962 (E. and I. Munroe) [CNC].
Texas: Hildalgo Co., Mercedes, 1 o’, Aug. 31, 1958 (H. Smalzried) [AMNH].
WASHINGTON: Chelan Co., Leavenworth, 2 o, July 3, 1949 (E. C. Johnston) [CNC]; Whitman Co., Pullman, 1 o, Aug. 6, 1898 (C. V. Piper) [USNM], from the paratype series of Bandera virginella, No. 11851 [USNM], genitalia slide, Carl Heinrich No. 6, Dec. 20, 1932.
CANADA: Britisn Cotumsia: Oliver, 1 9, Aug. 2, 1953 (D. F. Hardwick) [CNC]; 1 o&, Aug. 10, 1953; 1 @#, Aug. 11, 1953; 1 o, Sept. 14, 1953.
ALBERTA: Dominion Range Sta., Manyberries, 1 o, Aug. 14, 1951 (D. F. Hard- wick) [CNC].
Discussion.—The presumed habitat differences and the wide geo- graphic separation between eastern and western populations indicate that they may well represent distinct biological species, but the absence of clear-cut morphological differences between them dictates that better criteria must be found if a new species name is to be given to the eastern population. Eastern specimens differ in having the frons, vertex, occiput, tegula, and prothorax covered with brown scales. The forewing ground color is reddish brown rather than yellow, and the transverse posterior line of dots is usually better developed.
Bandera cupidinella Hulst
FIGuREs 32, 137, 172
Bandera cupidinella Hulst, 1888, p. 118. — Ragonot, 1889, p. 117. — Hulst, 1890, p. 203. — Smith, 1891, p. 84. — Ragonot, 1901, p. 410. — Hulst, 1902
72 U.S. NATIONAL MUSEUM BULLETIN 280
p. 440. — Dyar, 1908c, p. 116. — Barnes and McDunnough, 1917, p. 150. — Hampson, 1918, p. 90. — McDunnough, 1939, p. 36. — Rindge, 1955, p. 160. — Heinrich, 1956, pp. 315-316.
Anerastia conspersella Ragonot, 1901, p. 404. — Barnes and McDunnough, 1917, p. 149. — McDunnough, 1939, p. 36.
Rhinaphe conspersella (Ragonot). — Hampson, 1918, p. 86.
Nasutes venata Hampson, 1930, p. 54. — MecDunnough, 1939, p. 28 [listed in Phycitinae]. — Heinrich, 1956, pp. 315-316 [listed as a synonym of cupidinella].
Diacenosis.—This species may be recognized either by the absence of hair brushes in the male, or by the presence of a basal yellow margin on the costa of forewing of both sexes.
DescripTion.—Frons about as long as diameter of eye, light brown; lateral scales of labial palpi with brown restricted to a subterminal band; vertex, occiput, and thorax light yellow brown dorsally, occiput and prothorax gray brown laterally; abdomen varying light brown to brown.
Forewing with costa abruptly yellow marginally at base to one-fifth distance to apex; yellow to orange yellow between veins; discal cell, area between cell and vein As, and area posterior to vein Ag solid yellow with no brown scales; fringe on outer margin of alternating white and brown bands; brown beneath.
Male genitalia with transtilla complete, weakly sclerotized medially. Lateral knobs of juxta poorly developed. Vinculum bluntly rounded anteriorly. Aedeagus tapering very slightly posteriorly, bluntly rounded anteriorly. Eighth abdominal segment without lateral hair tufts.
Female genitalia with ovipositor setose. Ductus bursae broad.
Typrres.—B. cupidinella, in the U.S. National Museum; N. venata and A. conspersella, in the British Museum (Natural History).
Type pata.—B. cupidinella, lectotype male, hereby designated, Colorado, Denver (Fernald collection), USNM 40074, genitalia slide No. 575, J. Shaffer, Nov. 23, 1964.
N. venata, in the original description Hampson reports: Colorado, Boulder, 1 o&; Cockerell; alar expanse 20 mm. Type examined by Mr. Paul Whalley.
A. conspersella, lectotype male, hereby designated, Colorado, Akron, “11/9/83”; genitalia slide No. 705, J. Shaffer, Jan. 4, 1967.
SPECIMENS EXAMINED.—26 of, 2 9.
DistriBuTION (Map 4).
UNITED STATES: Cotorapo: Boulder Co., Boulder, 27, September 5 (Cocke- rell) [USNM]; Denver Co., Denver, 2 o, Sept. 1, 1904; 1 o&, Sept. 3, 1904;9 0, no date; Denver, 4 o, no date [USN M]; Prowers Co., Lamar, 4 co’, 1 9, Sept. 24,
1945 (B. C. Johnston) [CNC]; Pueblo Co., Pueblo, 1 o, September 1899 [AMNH]; no locality, 11 #@ [USNM].
REVISION OF PEORIINAE AND ANERASTIINAE (3
New Mexico: Hot Springs (7000 ft.), 1 o, September [USN M]; no locality (7000 ft.), 1 2, 1927 (H. S. Parish) [CU]. Texas: Jeff Davis Co., Ft. Davis (5000 ft.), 1 o&, 1927 (H. S. Parish) [CU].
Discussion.—Two males in the American Museum of Natural History are labeled ‘Type’ from Hot Springs, N. Mex., 7000 ft., August, September. A male genitalia slide of the August specimen was prepared by Carl Heinrich, No. 3226, June 14, 1946. The type locality is given as Colorado and these specimens must be regarded as another example of Hulst pseudotypes. The specimen in the U.S. National Museum labeled ‘“‘Type” is from Colorado and is assumed to be genuine.
Bandera virginella Dyar
FicuREs 33, 138, 173
Bandera virginella Dyar, 1908c, pp. 116-117. — Barnes and MeDunnough, 1917, p. 150. — MecDunnough, 1939, p. 36.
Diacnosis.—In the other members of this genus the discal cell and the area between the cell and vein A: are scaled with solid yellow, but in virginella these areas are pale yellow sprinkled with brown scales. The male genitalia are identical with those of cupidinella, but the eighth abdominal segment bears lateral hair tufts.
DescripTion.—Frons about two-thirds as long as diameter of eye, dirty white dorsally and light brown ventrally; tongue somewhat less well developed than on other members of the genus; vertex, occiput, and thorax light yellow brown dorsally, occiput and prothorax gray brown laterally; abdomen varying light brown to brown.
Forewings with areas between veins pale yellowish white, variously sprinkled with brown scales; fold of A, traced with brown scales; brown beneath.
Male genitalia as in B. cupidinella. Kighth abdominal segment with hair tufts similar to those of B. binotella, but with fewer scales and therefore smaller.
Female genitalia with ovipositor sparsely setose. Ductus bursae slender.
Typr.—In the U.S. National Museum.
Type pata.—Holotype: Washington, Pullman, male, July 24, 1898; C. V. Piper; Washington Exp. Sta. No. 463; USNM 11851; male genitalia slide No. 5, Carl Heinrich, Dec. 20, 1932.
Paratypes: Five specimens, Washington, Pullman, males; C. V. Piper; USNM 11851. Dates are July 8, 1898, Aug. 6, 1898, Aug. 10, 1898, July 31, 1899, Aug. 3, 1899. The specimen collected Aug. 6, 1898 bears genitalia slide label C.H. No. 6, Dec. 20, 1932 and belongs to B. binotella.
OTHER SPECIMENS EXAMINED.—51 o',7 @.
74. U.S. NATIONAL MUSEUM BULLETIN 280
DistriputTion (Map 4).—New Mexico to California, north to Alberta and British Columbia.
UNITED STATES: Arizona: Cochise Co., Southwestern Res. Sta., Chiri- cahua Mts., 1 o, Apr. 11, 1962 (Carl W. Kirkwood) [CNC]; 1 o&, Apr. 18, 1962; Yavapai Co., Prescott, 5 @, May 6, 1950 (E. C. Johnston) [CNC].
CALIFORNIA: Riverside Co., Kenworthy, 6 co’, June 9, 1937 (E. C. Johnston) [CNC]; Lake Hemet, 1 o&, June 9, 1937 (E. C. Johnston) [CNC]; San Bernardino Co., 12 mi. southeast of Ivanpan, 1 o&, May 1, 1956 (J. Powell) [UCB]; Upper Santa Ana River, 1 @, Aug. 12, 1948 (Grace H. and John L. Sperry) [AMNH]; Siskiyou Co., Indian Butte, 4 @, July 16, 1986 (E. C. Johnston) [CNC]; Ventura Co., Camp Ozena, Upper Cuyama, 4 o’, June 13, 1963 (C. W. Kirkwood) [CPK]; 3 oc’, June 14, 1963; 1 o&, June 18, 1963, 1 o, June 19, 1963; 2 o&, June 24, 1963; 1 o, June 27, 1963.
Couorapo: Rock Creek Canyon, 1 o, Sept. 26, 1957 (Margot May) [CNC]; 1 o, Sept.28, 1957.
New Mexico: Eddy Co., White City, 1 o&, May 15, 1950 (E. C. Johnston) [CNC].
WasHINGTON: Walla Walla Co., Walla Walla, 1 9, Apr. 9, 1931 (D. R. Bran- non) [USNM]; 1 o&, Apr. 21, 1931; 3 o, Apr. 18, 1935 (H. P. Lanchaster) ; Whitman Co., Pullman, 1 &, June 19, 1930 (J. F. Clarke) [USNM];1 ¢, 1 9, Aug. 3, 19382; 2 o#, July 3, 1935; Yakima Co., Satus Creek, 2 ¢, Aug. 19, 1949 (EH. C. Johnston) [CNC]; 1 o, Sept. 16, 1949; Tieton, 1 o&, May 12, 1931 (Fred Deari) [USNM].
CANADA: Axperta: Medicine Hat, 1 o, June 24, 1945 (K. Bowman) [CNC].
British Cotumsia: Shingle Cr. Road, Keremeos, 1 o, June 8, 1935 (A. N. Gartrell) [CNC]; 3 9, July 15, 1935; 2 9, July 22, 1935; Oliver (1000 ft.), 1 ¢, Aug. 10, 1953 (D. F. Hardwick) [CNC]; 1 o&, Aug. 11, 1953.
No data given except S. Diego, 1 o’, July 19, 1924 [USNM].
Wakulla, new genus
Typp.—Bandera carneella Barnes and McDunnough, 1918.
Diagnosis.—Among the phycitine genera exhibiting a reduced tongue this is the only genus in which the males have both hair tufts on the eighth abdominal segment and a spatulate uncus.
Description.—Frons conical, about two-thirds as long as eye diameter; labial palpi slightly ascending and extending somewhat beyond frons in male, female ascending, reaching beyond vertex; maxillary palpi well developed, cylindrical; tongue reduced, visible between bases of palpi, scaled at base; antennae filiform in both sexes, compressed, ciliate ventrally, cilia short, about one-sixth as long as segment width; ocelli minute.
Forewing and hindwing venation as in Bandera.
Male genitalia with uncus spatulate, setose dorsally and laterally. Gnathos arms each terminating in anterior hook; apical process bi- furcate. Transtilla incomplete. Juxta U-shaped, surrounding aedeagus ventrally and laterally. Vinculum stout, tapering, broadly truncate anteriorly, length and width about equal. Valvae with costa tubular,
REVISION OF PEORIINAE AND ANERASTIINAE 15
terminating in short free spine just before apex of cucullus; sacculus inflated on basal one-fourth of valve. Aedeagus stout, slightly tapering distally, vesica unarmed. Eighth abdominal segment of male with hair tufts.
Female genitalia with ovipositor tapering distally, sparsely setose, about 114 times as long as broad. Apophyses well sclerotized, posterior about four to five times as long as basal separation, anterior slightly less than twice as long as basal separation. Ductus bursae long. Bursa small, bearing three small barlike signae. Ductus seminalis leaving bursa near posterior end, just posterior to signae, threadlike, well sclerotized.
Wakulla carneella (Barnes and McDunnough), new combination Fiauress 43, 100, 139, 174
Bandera carneella Barnes and McDunnough, 1913, p. 184; 1917, p. 150. — Gross- beck, 1917, p. 1384. — McDunnough, 1939, p. 36. — Kimball, 1965, p. 251.
DriaGnosis.—This is the only known species in the genus.
Description.—Labial palpi with basal segments white, second and third segments light brown or reddish brown on outer sides, white or nearly so on inner sides; maxillary palpi reddish brown; antennae with scape brown, shaft light brown; frons, vertex, occiput, patagia, and tegulae reddish brown.
Forewing reddish brown, sprinkled with dark brown scales; two small dark brown spots at distal end of cell; male with tuft of black scales at base of costa. Hindwing light brown.
Genitalia as described for the genus.
Typr.—tIn the U.S. National Museum.
Type pata.—Holotype male, Everglades Fla., Apr. 8-15, Barnes collection, “Photograph pl. 1 No. 5”; USNM 69384; genitalia slide No. 574, J. Shaffer, Nov. 23, 1965.
SPECIMENS EXAMINED.—3 0, 19.
DistrisuTion (Map 3).—Known only from Florida.
UNITED STATES: Fioripa; Manatee Co., Gulf Coast Exp. Sta., Bradenton,
14, Aug. 5, 1955 (E. G. Kelsheimer) [CPK]; 1¢, Mar. 28, 1956; Sarasota Co., Siesta Key, 1c, Feb. 13, 1951 (C. P. Kimball) [CPK]; 19, Nov. 9, 1952.
Tampa Ragonot
Tampa Ragonot, 1887, pp. 19-20; 1889, p. 117. — Hulst, 1890, p. 203 [listed in Phycitinae]. — Smith, 1891, p. 84 [listed in Phycitinae]. — Ragonot, 1901, p. 411. — Hulst, 1902, p. 440. — Barnes and McDunnough, 1917, p. 150. — Forbes, 1923, p. 637. — Hampson, 1918, p. 62. — McDunnough, 1939, p. 36. (Type: Tampa dimediatella, 1887. Monobasic.]
285—-934—68—6
76 U.S. NATIONAL MUSEUM BULLETIN 280
Dracnosis.—The presence of only two radial veins in the forewing distinguishes Tampa among the phycitine genera showing a reduced tongue.
Description.—Frons conical; labial palpi ascending, often porrect in females, about three times eye diameter in males, 3% times in females; maxillary palpi three-segmented, cylindrical, reaching frons or nearly so; tongue reduced, usually visible between palpi; antennae filiform in both sexes, somewhat thicker in male, female scaled dor- sally and laterally, ventrally with short appressed cilia, male scaled dorsally, ventrally with cilia about one-half segment width, hooked at tips; ocelli absent.
Forewing with 8 veins; R, from near upper outer angle of cell, R244 fused, from the angle; M, from about one-third from top of cell; M,,; fused, from lower outer angle; Cu; from just before the angle; Cu, from well before the angle. Hindwing with 7 veins; Sc and Rs long stalked: M, from upper outer angle; M2,3 fused, long stalked with Cu,, from lower outer angle; Cu, from just before the angle.
Male genitalia with uncus tapering, broadly and bluntly rounded apically; rather densely setose dorsally and laterally. Gnathos with medial process small, simple, shallowly notched apically. Transtilla complete, weakly sclerotized. Juxta crescent shaped, with pair of lateral setose tubercles. Vinculum V-shaped, apex bluntly rounded. Valve narrow, sides nearly parallel, apex rounded. Vesica unarmed.
Female genitalia with ovipositor tapering distally, about twice as long as wide. Apophyses slender, posterior about six or seven times as long as their basal separation; anterior about 2); times as long as their basal separation, diverging. Ductus bursae long, slender. Bursa small, with a group of five to seven parallel barlike signae extending anteriorly from ductus seminalis near posterior end. Ductus seminalis from posterior end of bursa, tapering, then slender.
Tampa dimediatella Ragonot Ficurss 35, 69, 101, 140, 175
Tampa dimediatella Ragonot, 1887, p. 20; 1889, p. 117. — Hulst, 1890, p. 203. — Smith, 1891, p. 84. — Ragonot, 1901, p. 411. — Hulst, 1902, p. 440. — Barnes and McDunnough, 1917, p. 150. — Grossbeck, 1917, p. 134. — Hampson, 1918, p. 63. — Forbes, 1923, p. 367. — McDunnough, 1939, p. 36. — Kimball, 1965, p. 251.
Dracnosis.—This is the only known species in the genus.
Description.—Labial palpi with basal segments and ventral and inner sides of second segments white, third segments and outer sides of second dark brown to black; maxillary palpi, frons, vertex, sides of occiput dark brown to black; dorsum of occiput, patagia, and
REVISION OF PEORIINAE AND ANERASTIINAE “a
tegulae somewhat lighter; antennae brown; legs white on inner sides, outer sides varying light brown to dark brown.
Forewings light brown anterior to cell, sprinkled with dark brown scales; cell a white band extending to wing apex, sprinkled with a few dark brown scales; discal spot small, black; cubitus traced with black sometimes extending from cell to A; fold; wings grayish-red posterior to black trace; As traced with black. Underside brown to grayish brown, cell lighter. Hindwings light brown above and below, darker apically.
Genitalia as described for the genus.
Typr.—tIn the Muséum National d’Histoire Naturelle.
Typr pATA.—Lectotype male, hereby disignated, Archer, Florida, "23/3 82”: collection C. V. Riley; male genitalia slide No. 550, J. Shaffer, July 28, 1965.
SPECIMENS EXAMINED.—180 o’, 39 @.
DistriputTion (Map 1).—Atlantic and Gulf Coastal Plain, eastern Texas to Florida, northward